Plants terrestrial, 10-140 cm tall. Rhizome short, erect to ascending, densely scaly at apex and at stipe bases. Fronds clustered; stipe dark brown to stramineous, scaly throughout; scales of two different types: (1) lanceolate, membranous, appressed, and iridescent scales, and (2) subulate, stiff, spreading scales without obvious iridescence. Lamina oblong-lanceolate to ovate-triangular, broadest at base, 1-4-pinnate; basal pair of pinnae always largest, triangular or broadly lanceolate, basal basiscopic pinnules of basal pinnae usually much enlarged; distal pinnae or pinnules ± adnate to rachis or costae and ± decurrent at their bases; veins all free, veinlets on ultimate pinnules or lobes simple or (rarely) bifurcate to pinnate; lamina texture herbaceous or papery, rarely leathery, with ctenitoid hairs on adaxial surface and short glandular hairs on abaxial surface, or rarely glabrous; costae of pinnules prominent, always covered with ctenitoid (multicellular) hairs. Sori medial or rarely submarginal; indusia present or not, sometimes very small and hidden by maturing sporangia. Perispore echinate, coarsely folded, or tuberculate. x = 41.
Plants generally terrestrial. Stems erect to obliquely ascending, stolons absent. Leaves monomorphic, evergreen or dying back in winter. Petiole 2/3 to equaling length of blade, base not swollen; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade lanceolate to deltate, 1--4-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous. Pinnae not articulate to rachis, segment margins nearly entire to crenulate, ciliate; proximal pinnae not reduced, sometimes basal pair much the longest, sessile to petiolulate, equilateral or inequilateral with basiscopic side more developed (pinnules noticeably longer); costae adaxially rounded or flat, not grooved; indument of linear to lanceolate scales and often multicellular glandular hairs abaxially, of multicellular reddish hairs adaxially. Veins free, simple or forked. Sori in 1 row between midrib and margin, round; indusia round-reniform, attached at narrow sinus, sometimes small or seemingly absent, persistent or caducous. Spores brownish, usually spiny, sometimes prominently cristate, rarely finely reticulate. x = 41.
Sori dorsal on veins; indusium round-reniform, sts absent, sporangia stalked; annulus incomplete, vertical. Spores bilateral, echinulate. Rhizome short, erect, or creeping, paleate; stipes paleate. Lamina bipinnate to decompound, catadromic; hairs pluricellular, constricted between cells. Terrestrial ferns of about 150 widely distributed spp.
Fronds usually much longer than wide; basiscopic margin of lamina-lobes not thickened.