Ovary 2-locular; ovules numerous on placentas attached to the septum; style usually with pollen collecting hairs at the level of the anthers; stigma/pollen presenter with 2 linear lobes, exserted or mostly included.
Calyx tube ellipsoid or obconic; limb-tube ± obsolete or only evident in fruit; lobes contorted, foliaceous, ± equal, mostly erect during flowering, venose, persistent.
Corolla tube long, narrowly cylindrical; lobes linear or linear-lanceolate to elliptic, usually long and pointed, spreading, contorted in the bud.
Flowers mostly rather large, hermaphrodite, 4–6-merous in few-to many-flowered cymes at the extremities of terminal shoots and lateral branches.
Leaves petiolate, opposite, sometimes slightly unequal in size, usually with distinct pubescent domatia; stipules usually conspicuous.
Shrubs or small trees, nearly always erect, rarely slightly scrambling or one species a prostrate subshrub.
Fruit subglobose or oblong to oblong-conic, scarcely or slightly fleshy, often longitudinally ribbed.
Stamens sessile, inserted below the level of the throat, included or anther apices exserted.
Seeds numerous, small, mostly angular, smooth and glossy.