Lithospermum L.

Stoneseed (en), Grémil (fr)

Genus

Angiosperms > Boraginales > Boraginaceae

Characteristics

Perennial or rarely annual or biennial erect or spreading herbs or subshrubs with hispid, strigose or densely hairy stems. Leaves alternate, entire, the venation usually obscure. Flowers yellow, orange or white, regular, sometimes heterostylous, in few–several-flowered simple or paired, terminal or axillary scorpioid cymes; bracts leafy, numerous, exceeding the calyx. Calyx-lobes 5, linear-cuneate or lanceolate, usually much shorter than the corolla-tube, the abaxial the largest. Corolla funnel-shaped or cylindrical, velvety outside; tube straight, glabrous or pubescent at the base inside, the throat with small velvety or glandular appendages, less often naked; lobes 5, spreading or rarely erect, imbricate, rounded, ovate, obovate or semicircular; nectaries on the internal face and at the base of the tube either as swellings, a narrow ring, lobes or small hairs. Stamens inserted near the middle of the tube, included. Ovary with 4 lobes; style gynobasic, simple, slender to rather stout, included or exserted; stigmas 2, globose, hemispherical or ellipsoid, terminal or situated just below the bilobed apex of the style. Fruit of 4 nutlets or less by abortion, ovoid or ellipsoid, mostly very shiny and porcellanous, rarely verrucose or rugose.
More
Herbs annual or perennial, short strigose. Leaves alternate. Cymes terminal or flowers solitary, bracteate. Calyx 5-parted nearly to base, slightly enlarged in fruit. Corolla white, yellow, or violet, actinomorphic, funnelform or salverform; throat with appendages or bands of hairs, or longitudinally crispate; limb usually campanulate, 5-parted; lobes spreading. Stamens included; filament very short; anthers oblong-linear, apex obtuse, mucronulate. Style filiform, not exserted; stigma entire or indistinctly 2-cleft, capitate. Gynobase flat. Nutlets white or gray, ovoid, smooth, shiny or tuberculate; attachment scar at base adaxially.
Cor funnelform or salverform, with or without fornices; anthers included or partly exserted; gynobase low-pyramidal or flat or depressed; nutlets smooth to pitted or wrinkled, basally attached, the large scar often surrounded by a sharp rim, sometimes only one nutlet maturing; annual to more often perennial herbs, seldom pungently hairy, with mostly yellow or white or greenish-white fls in modified leafy-bracteate cymes, or solitary in or near the upper axils, often heterostylic; fruiting pedicels mostly erect or ascending. (Buglossoides) 75, widespread, mostly temperate and mountainous regions.
Annual, biennial or perennial herbs, sometimes shrubs, hispid or with soft hairs. Infl. short, strongly bracteate, cymose, often terminal. Calyx 5-lobed, sometimes divided to base, persistent. Corolla cylindric, usually funnelform, rarely salverform; tube > lobes, with 5 longitudinal folds; lobes white, yellowish, blue or purple; throat with bands of hairs or scales. Stamens included. Stigma capitate or 2-lobed. Nutlets smooth or tuberculate, shining, very hard.
Fruit dry, consisting of 4 nutlets or less by abortion; nutlets detaching completely from the receptacle, ovoid to ellipsoid, erect, smooth, white and shining, with the dorsal surface convex and the ventral one obtusely angular with a flat or convex point of attachment.
Corolla cylindrical to infundibuliform, orange, yellowish or whitish; tube with 5 more or less glandular invaginations in the throat and with a ring of nectaries at the base, inside; lobes imbricate in bud.
Style gynobasic, filiform, entire or sometimes slightly 2-fid at the apex, included or sometimes almost exserted.
Flowers 5-merous, actinomorphic, isostylous or heterostylous, on leafy terminal or axillary cymes.
Stamens inserted about the middle or near the top of the corolla tube, included.
Ovary with 4 free lobes, inserted on the narrow gynobase.
Stigmas 2, terminal or subtenninal or 1, bilobed.
Perennial herbs or undershrubs.
Calyx lobed to near the base.
Leaves alternate, entire.
Life form perennial
Growth form herb
Growth support -
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.1 - 0.3
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-9

Usage

Uses medicinal
Edible -
Therapeutic use Antidote (unspecified), Febrifuge (unspecified), Antiphlogistic (unspecified), Granulation (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Lithospermum unspecified picture

Distribution

Lithospermum world distribution map, present in China, France, New Zealand, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:325901-2
WFO ID wfo-4000021988
COL ID 5FPB
BDTFX ID 86718
INPN ID 194220
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lithospermum Macromeria Onosmodium Cyphorima Lasiarrhenum Nomosa Osmodium Pentalophus Psilolaemus

Lower taxons

Lithospermum macbridei Lithospermum mediale Lithospermum cinerascens Lithospermum erythrorhizon Lithospermum gayanum Lithospermum cuzcoense Lithospermum leymebambense Lithospermum rodriguezii Lithospermum bolivariense Lithospermum azuayense Lithospermum chiapense Lithospermum dodrantale Lithospermum exsertum Lithospermum helleri Lithospermum johnstonii Lithospermum leonotis Lithospermum macromeria Lithospermum notatum Lithospermum oaxacanum Lithospermum decipiens Lithospermum parviflorum Lithospermum pinetorum Lithospermum rosei Lithospermum rzedowskii Lithospermum trinervium Lithospermum turneri Lithospermum unicum Lithospermum canescens Lithospermum calcicola Lithospermum discolor Lithospermum matamorense Lithospermum oblongifolium Lithospermum strictum Lithospermum californicum Lithospermum calycosum Lithospermum caroliniense Lithospermum cobrense Lithospermum confine Lithospermum latifolium Lithospermum mirabile Lithospermum multiflorum Lithospermum ruderale Lithospermum tuberosum Lithospermum viride Lithospermum affine Lithospermum cinereum Lithospermum diversifolium Lithospermum flexuosum Lithospermum hirsutum Lithospermum papillosum Lithospermum scabrum Lithospermum distichum Lithospermum guatemalense Lithospermum cuneifolium Lithospermum pringlei Lithospermum revolutum Lithospermum jimulcense Lithospermum indecorum Lithospermum flavum Lithospermum berlandieri Lithospermum virginianum Lithospermum tubuliflorum Lithospermum nelsonii Lithospermum rosmarinifolium Lithospermum obovatum Lithospermum ireneae Lithospermum muelleri Lithospermum peruvianum Lithospermum occidentale Lithospermum subsetosum Lithospermum molle Lithospermum album Lithospermum barbigerum Lithospermum incisum Lithospermum officinale Lithospermum afromontanum