Spiranthes Rich.

Ladies'-tresses (en), Spiranthe (fr)

Genus

Angiosperms > Asparagales > Orchidaceae

Characteristics

Sep and lateral pet similar, the pet connivent with the upper sep (or with all the sep) and projecting forward over the lip and column; lip oblong or ovate to pandurate in outline, bearing 2 small callosities near the basal angles, its margins below the middle upturned and embracing the short column, the more distal part ± decurved, not crested or ridged, but often with a median area of different color; anther borne on the back of the column, its 4 pollinia attached to the viscidium and commonly surpassed by the bidentate or bifid rostellar beak; slender herbs with narrow (seldom broader), chiefly basal lvs, the erect stem bearing several reduced lvs (or mere sheaths) and a terminal, spirally twisted, spike-like raceme; fls mostly white or whitish at least in large part. (Ibidium) 300, cosmop. The fls decrease in size progressively from the bottom to the top of the infl. Measurements given here are for fls in the lower third. In spp. 1–5 the infl is ordinarily such a close spiral that the fls appear to be in 3 or 4 vertical, straight or somewhat spiral ranks. In spp. 6–11 the infl is typically more lax and open, with the fls obviously in a single long spiral, but some of these spp. vary to forms with the infl approaching that of the first group. A number of hybrids have been documented.
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Herbs, terrestrial, acaulescent. Roots fasciculate, fusiform, fleshy, glabrous. Leaves basal, forming a rosette, linear, elliptic, or broadly ovate, rarely subcylindric, base sheathing. Inflorescence terminal, racemose with many small flowers arranged spirally around rachis. Flowers resupinate, horizontal or nodding, not opening widely, small; ovary twisted, fusiform. Sepals free, narrowly elliptic to lanceolate, subsimilar; dorsal sepal erect, often connivent with petals and forming a hood; lateral sepals dilated or sometimes saccate at base. Petals erect, oblanceolate, recurved; lip entire or obscurely 3-lobed, shortly clawed, apex recurved, undulate, lateral margins embracing column; disk usually papillose, with 2 fleshy glands at base. Column clavate, ventrally pubescent; anther erect, 2-locular, on dorsal surface of column; pollinia 2, each 2-partite, granular-farinaceous, with or without short caudicle and attached to narrow viscidium; rostellum erect, 2-lobed at apex after removal of viscidium; stigma suborbicular to elliptic. Capsule ellipsoid.
Small, or rarely large, terrestrial (rarely epiphytic) herbs with basal or cauline leaves or both, or leafless. Roots often fleshy, fasciculated or tuberous. Leaves various, produced before, with, or after, the flowers. Sepals free; dorsal sepal usually erect and forming a galea with the petals; lateral ones erect or spreading, affixed to the summit of the ovary, decurrent and forming a free or adnate mentum. Petals usually narrow and usually coherent to the dorsal sepal. Lip sessile or clawed, plane, concave or gibbous, simple or lobed, in some species bi-caudate at the base, adherent to the column in almost all the species, ecallose or callose. Column terete; clinandrium often membranaceous and conspicuous, often continued into the rostellum; rostellum various, inconspicuous or conspicuous, truncate and retuse to lobed to aristate; anther dorsal, erect, sessile or stipitate; pollinia 2, powdery or granular, usually attenuated at one end.
Fls ∞ in ± spirally twisted spike; floral bracts = or > ovary. Ovary and per. ± glandular-pubescent; per. ± horizontal-tubular, dorsal sepal uppermost; lateral sepals subsimilar, not connate; petals ± similar, in part adnate to dorsal sepal. Labellum of about equal length, sessile or shortly clawed, concave and embracing base of column, distally spreading and crenate to crisped; calli 2, lateral, near base. Column short, us. flanked by 2 small wings; anther dorsal, often overtopped by rostellum, pollinia 2 per cell, pollen granular; stigma broad; rostellum us. long, remaining as 2 tall processes after its central long-oval viscidium is detached. Plants terrestrial, glab. except for infl.; roots clustered, us. ± tuberous. Lvs several, mostly basal; stem with persistent sheathing bracts Spp. estimated at 50 to 300, widely dispersed through temperate zones. The N.Z. sp. widespread in Australia and Asia.
Herbs, perennial, terrestrial. Roots fleshy, fasciculate, slender to tuberous. Stems with foliaceous sheaths. Leaves basal or occasionally cauline. Inflorescences terminal spikes, flowers inserted in lax to dense, ± evident spiral, pubescent to glabrous, often glandular. Flowers resupinate, often somewhat apically recurved, white, cream, or yellow (pink in S. sinensis); perianth parts distinct or sepals basally connate, connivent; lip lanceolate to ovate or pandurate, rarely linear, fleshy to somewhat membranaceous, base with pair of calli, apex usually ± crisped; mentum or spur absent; column short, cylindric; anther cordate, apex acute or obtuse; pollinia clavate with slender viscidium; ovary sessile, cylindric.
Terrestrial, evergreen or facultatively deciduous orchids with thick, fleshy roots. Leaves smooth, narrow, in a loose basal rosette. Inforescence tall, wiry, spicate. Flowers small, colourful, densely crowded in a spiral spike. Basal part of dorsal sepal and petals overlap loosely to form a galea which partially hoods the labellum. Lateral sepals free, porrect to spreading. Labellum flexibly attached to anterior base of column. Labellum lamina 3-lobed, protruding shortly from flower; base tubular, with 2 rounded or tonsil-like glands; distal and apical margins wavy to frilly. Column slender.
Life form perennial
Growth form herb
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Foliage retention evergreen
Sexuality hermaphrodite
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Environment

These orchids grow in moist to wet soil in open, grassy habitats.
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Hardiness (USDA) 5-11

Usage

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Cultivation

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