Theobroma cacao L.

Cacao (en), cacaoyer (fr)

Species

Angiosperms > Malvales > Malvaceae > Theobroma

Characteristics

Tree usually 4-8 m. high, rarely taller and up to 20 m.; growth below jorquette; primary branching quinate; young branchlets terete, grayish-green or brownish, densely or sparsely pubescent with slender, patulous, acute, simple or furcate hairs, later glabrate, rugulose and sparsely lenticellate; stipules subulate, deciduous. Leaves alternate, distichous, the petioles thickened-pulvinate at both ends, 1.5-2 (1-3) cm. long, on orthotropous stems 3-10 cm. long, pubescent or tomentose with slender, spreading, acute, simple hairs; blade subobovate-oblong or elliptic-oblong, slightly asymmetrical, rounded or obtuse at base, attenuate and acuminate at apex, entire or slightly and irregularly sinuate, 15-50 cm. long and 4-15 cm. broad, cori-aceous or chartaceous, the acumen acute or subacute and 1-2.5 mm. long; upper surface green, glabrous except for the pubescent or puberulous, prominent costa, the secondary nerves filiform; lower surface lighter green, glabrous or with very sparse, minute, simple, furcate or stellate hairs, rarely puberulous, the costa prominent, the 9-12 secondary nerves on each side prominent, subpatulous. Infiorescences on the trunk and on the branches, usually borne on small tubercles, the cymose peduncles 1-3 mm. long, stellate-pubescent, hirtellous and with scattered, glandular hairs; bracts and bracteoles ovate or ovate-oblong, acute or subacute, pubescent, deciduous; pedicels capillary, rigid, 5-15 mm. long, pale green, whitish or reddish, hirtellous with patulous, stellate or furcate hairs and glandular, capitate hairs; buds ovoid or oblong-ovoid, acute, white, whitish-green, lilac or reddish, subglabrous or sparsely puberulous. Flowers with lanceolate or oblong-lanceolate sepals, these shortly united at base, acute at apex, 5-8 mm. long and 1.5-2 mm. wide, thick-membranaceous, white, greenish-white, pale violaceous or reddish, slightly 3-nerved, glabrous or with stellate and glandular hairs outside, glabrous or with rare, glan-dular hairs inside; petals thick-membranaceous; petal-hoods obovate, rounded at apex, 3-4 mm. long and 5-2 mm. wide, white, 3-nerved; petal-laminae stipitate, the stipe linear and 2-4 mm. long, obovate or rombic-obovate, the apex attenuate, acuminate or subtruncate and shortly mucronate, rarely blunt, the margin entire or sinuate, 1.5-2.5 mm. long and 1.5-2 mm. wide, pale yellowish; staminodes nar-rowly subulate, very acute, 4-6 mm. long, the middle vein thick, angular, red or purplish, minutely papillose-pilose, the thin margin whitish, ciliate with slender, flexuose, simple hairs; filaments 1.5-2 mm. long, glabrous, 2-antheriferous; ovary glabrous or usually glandular, the glands numerous, stipitate, white or reddish, pluricellular; styles adherent, glabrous. Fruit variable in shape, from globose to fusiform and acute, with a very smooth to a strongly ridged and rugose or verrucose surface; pericarp consistently fleshy, 5-15 mm. thick, usually made of 2 more or less conspicuously different, carnose layers (epicarp and endocarp) separated by a thin, ligneous membrane (mesocarp), the latter sometimes reduced to isolated bundles of fibers or lacking, the endocarp also sometimes lacking; pulp white and sweet; seeds 20-40, ovoid, ellipsoid, amygdaloid, more or less complanate or almost round in cross-section, variable in size, 20-40 mm. long and 12-20 mm. broad, the testa subcoriaceous, brown; cotyledons white, purplish, violet or intermediate in color.
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Evergreen trees, to 12 m tall; bark thick, dark gray-brown. Branchlets brown, puberulent. Stipules linear, caducous; leaf blade narrowly ovate-to obovate-elliptic, 20-30 × 7-10 cm, both surfaces glabrous or sparsely stellate, base rounded to shallowly cordate, apex long acuminate. Inflorescence small and delicate, cymose. Flowers ca. 18 mm in diam.; pedicels ca. 12 mm. Calyx pink, lobes narrowly lanceolate, persistent, margin hairy. Petals 5, yellowish, lightly longer than calyx, lower part helmet-shaped and abruptly narrowed, reflexed, apex acute. Staminodes linear. Ovary obovoid, slightly 5-angular, 5-celled; ovules 14-16 per locule, in 2 rows; style cylindrical. Drupe ellipsoid or narrowly ellipsoid, 15-20 × ca. 7 cm, longitudinally 10-grooved; endocarp thick, fleshy, hard and woody when dried, 4-8 mm thick. Seeds 12-14 per cell, ovoid, slightly flattened, ca. 2.5 × 1.5 cm. Fl. throughout year.
A small evergreen tree. It grows to 15 m tall. Often it is pruned to be only 6 m tall. The trunk is straight for about 90-159 cm above the ground then a fan arrangement of branches develops. The leaves are large and simple. They are in a spiral arrangement around the branch. They are 17-30 cm long and hang down. They are red brown when young. The leaves are wider towards the pointed tip. The leaves are thin but leathery with prominent veins. The flowers are on short shoots like cushions on the trunk and woody branches. The flowers are small, yellow-white and about 1 cm across. A ribbed pod develops. The pods are large about 15-20 cm long by 10 cm across. They are green when young but become yellow or red. The pods have rows of seeds. There are 20-50 oval seeds. These are white at first then become red-brown as they ripen. There is a sweet pulp surrounding the seeds.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 10.0
Root system -
Rooting depth (meter) -
Root diameter (meter) 0.6
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

A tropical plant. It is native from Mexico to Central America. It grows in lowland areas with a rainfall around 2,000 mm annually. It can grow in areas with rainfall between 1,200 and 7,000 mm. Plants do best in acid or neutral soil which is well drained. It has a long tap root which will not develop on rocky or hard soil. Cocoa does not do well on windy sites where there are sudden fluctuations of humidity. Shade trees help provide these conditions. It needs temperatures above 20°C. In PNG it mostly grows below 800 m above sea level. It suits hardiness zones 11-12. In Yunnan.
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An understorey plant of evergreen rainforest in the wet humid tropics, growing in places that are not usually seasonally inundated.
Light 1-9
Soil humidity 4-6
Soil texture 1-4
Soil acidity -
Soil nutriment -
Hardiness (USDA) 10-12

Usage

The seeds are eaten. The seeds are also processed and used for chocolate. The seeds are fermented for 6 days to remove the flesh then washed and dried. The pulp in the pod is edible when ripe. It is used in jellies and sweets.
Uses afforestation animal food beverage charcoal environmental use fiber fodder food gene source material medicinal poison social use wood
Edible fruits seeds
Therapeutic use Anesthesia (leaf), Central nervous system stimulants (leaf), Constipation (seed), Diabetes mellitus (seed), Diet, food, and nutrition (seed), Diuretics (seed), Emollients (seed), Fissure in ano (seed), Hemorrhoids (seed), Migraine disorders (seed), Skin diseases (seed), Thrombophlebitis (seed), Fissure of nipple (seed), Antiseptic (unspecified), Burn (unspecified), Coffee (unspecified), Cough (unspecified), Dentifrice (unspecified), Dry-Lip (unspecified), Emmenagogue (unspecified), Eye (unspecified), Kidney (unspecified), Listlessness (unspecified), Parasiticide (unspecified), Parturition (unspecified), Pilatory (unspecified), Rheumatism (unspecified), Wound (unspecified), Bite(Snake) (unspecified), Diuretic (unspecified), Liqueur (unspecified), Pregnancy (unspecified), Suppository-Base (unspecified), Poison (unspecified), Alopecia (unspecified), Astringents (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Plants are grown from seed. Seed can be sown in pots in a nursery then transplanted or can be sown directly. Seedlings need shade. Plants can also be grown from cuttings. Cuttings from side branches are used. Cuttings are normally raised in a nursery. Seeds should be removed from the pod and dried but should not be stored for more than 3 months. Seeds germinate in 1 week. Plants should be spaced 3 m apart. Plants can be pruned. Flowers are pollinated by insects.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) 21 - 32
Size -
Vigor -
Productivity -

Images

Leaf

Theobroma cacao leaf picture by Daniel Barthelemy (cc-by-sa)
Theobroma cacao leaf picture by Pascal SIMON (cc-by-sa)
Theobroma cacao leaf picture by Thomas T.Akoto (cc-by-sa)

Flower

Theobroma cacao flower picture by Daniel Nossa-Silva (cc-by-sa)
Theobroma cacao flower picture by Dieter Albrecht (cc-by-sa)
Theobroma cacao flower picture by Dieter Albrecht (cc-by-sa)

Fruit

Theobroma cacao fruit picture by amila gunesekera (cc-by-sa)
Theobroma cacao fruit picture by Daniel Nossa-Silva (cc-by-sa)
Theobroma cacao fruit picture by Daniel Barthelemy (cc-by-sa)

Distribution

Theobroma cacao world distribution map, present in American Samoa, Antigua and Barbuda, Benin, Bonaire, Sint Eustatius and Saba, Belize, Brazil, Barbados, Central African Republic, China, Côte d'Ivoire, Cameroon, Cook Islands, Colombia, Costa Rica, Cuba, Cayman Islands, Dominica, Ecuador, Fiji, Micronesia (Federated States of), Gabon, Ghana, Guinea, Guadeloupe, Equatorial Guinea, Grenada, Guam, Honduras, India, Jamaica, Lao People's Democratic Republic, Liberia, Saint Lucia, Sri Lanka, Myanmar, Northern Mariana Islands, Martinique, Nigeria, Nicaragua, Niue, Panama, Philippines, Puerto Rico, Sierra Leone, Sao Tome and Principe, Thailand, Taiwan, Province of China, United States of America, Venezuela (Bolivarian Republic of), and Viet Nam

Identifiers

LSID urn:lsid:ipni.org:names:320783-2
WFO ID wfo-0000458440
COL ID 56BND
BDTFX ID -
INPN ID 447630
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Theobroma leiocarpum Theobroma sapidum Cacao sativa Cacao minar Theobroma sativum Theobroma kalagua Theobroma pentagonum Theobroma integerrimum Theobroma saltzmanniana Theobroma sativa Theobroma sphaerocarpum Theobroma salzmanniana Cacao minus Cacao theobroma Theobroma cacao subsp. leiocarpum Theobroma cacao var. leiocarpum Theobroma cacao subsp. sphaerocarpum Theobroma cacao f. leiocarpum Theobroma cacao subsp. cacao Theobroma sativa var. leucosperma Theobroma sativa var. melanosperma Theobroma cacao subsp. sativa Theobroma cacao subsp. pentagonum Theobroma cacao

Lower taxons

Theobroma cacao 'Caeri 1' Theobroma cacao 'Caeri 2' Theobroma cacao 'ICS 45' Theobroma cacao 'RIM 52' Theobroma cacao 'SPA 10' Theobroma cacao 'TARS 1' Theobroma cacao 'TARS 14' Theobroma cacao 'TARS 23' Theobroma cacao 'TARS 27' Theobroma cacao 'TARS 30' Theobroma cacao 'TARS 34' Theobroma cacao 'TARS 9'