Shrubs or small trees, some with lignotubers; branches erect or prostrate; bark smooth, rarely warty. Leaves simple, entire or lobed, or deeply segmented, glandular, sclerified. Inflorescence solitary, axillary or terminal, single-flowered. Flowers zygomorphic, bisexual, sessile, within an involucre of numerous, scale-like bracts. Perianth tubular, with 4 tepals, hirsute outside, bearded inside behind anthers. Stamen filaments fused to tepal claws; all stamens perfect and fertile, or adaxial stamen sterile. Hypogynous glands 4, alternating with tepals. Ovary sessile; style terminal; pollen presenter conical or flattened and elliptic; ovule 1. Fruit an achene.