Shrubs or small trees, evergreen or deciduous, perennial, monoecious or dioecious; stems and foliage without latex. Indumentum of simple, multicellular trichomes, glandular trichomes absent, stinging trichomes absent. Stipules entire, inconspicuous, deciduous. Leaves alternate, petiolate, elobate, penninerved, crenate or serrate, with glands at base of lamina or near apex of petiole. Inflorescences terminal or axillary, racemose, solitary, usually bisexual, with flowers in bracteate glomerules. Male flowers pedicellate: calyx lobes 2–4, valvate, free and ±equal; petals absent; disc of numerous glands; stamens 10–200, filaments free and attached to a slightly convex receptacle; anthers basifixed, bilobate, thecae globose to oblong and longitudinally dehiscent; pistillodes absent. Female flowers pedicellate: calyx lobes 3, valvate, shortly connate; petals absent; disc hypogynous, annular; ovary 2–4-locular, ovules uniloculate; styles shortly connate at base, simple and spreading. Fruit capsular, trilobate, surface smooth, dehiscing septicidally into 3 bivalved cocci. Seeds globose; testa crustaceous; albumen fleshy; ecarunculate; cotyledons broad, flat.
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Shrubs or trees, dioecious, rarely monoecious. Branchlets puberulent. Leaves alternate; stipules small or minute, deciduous; leaf blade mostly ± dentate; venation pinnate. Inflorescences axillary, unbranched; bracts small. Male flowers 1 to many per bract; sepals (2 or)3 or 4, valvate, closed in bud; petals absent; stamens (10-)20-30(-200); filaments free; anthers basifixed, 2-locular, theca almost free, erect; juxtastaminal glands many, short, erect, apex pilose; pistillode absent. Female flowers mostly 1 per bract; sepals 2-4; petals absent; disk annular or dissected; ovary 2-or 3(or 4)-locular; ovule 1 per locule; styles short, stigmatic surface papillose or plumose, recurved. Fruit a capsule, 2-or 3(or 4)-lobed. Seeds globose; testa fleshy, inner layer hard, foveolate-reticulate.
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