Shrubs or herbs, with cystoliths, isophyllous or slightly anisophyllous. Leaves petiolate. Inflorescences terminal and/or sometimes axillary, open panicles or thyrses; bracts small, shorter than calyx. Calyx deeply 5-lobed, lobes similar. Corolla funnel-shaped, 2-lipped; lower lip 3-lobed; upper lip shortly 2-lobed, lacking a stylar furrow (i.e., erugulate); lobes ascending cochlear in bud. Stamens 2; anthers 2-thecous; thecae equal or unequal, parallel (to perpendicular), subequally to unequally inserted, base muticous; staminodes absent; pollen 2-pororate [elsewhere also 3-colporate and 6-pseudocolpate]. Ovary with 2 ovules per locule. Capsule clavate with a solid sterile basal stalk, 4-seeded; retinacula present; septa with attached retinacula remaining attached to inner wall of mature capsule. Seeds subdiscoid.
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