Pittosporum Banks ex Gaertn.

Cheesewood (en)

Genus

Angiosperms > Apiales > Pittosporaceae

Characteristics

In Australia and its territories, erect shrubs, or more usually trees, to 30 m high and 45 cm diam. d.b.h., heteroblastic, most functionally dioecious but never completely so. Short shoots present and sometimes spinescent (e.g. Pittosporum oreillyanum); hairs T-shaped or uniseriate, clear, white, red-purple or golden brown. Leaves variable depending on growth stage. Cotyledons mostly 2, opposite. Juvenile leaves with lamina margin variably toothed or lobed, rarely pungent (P. multiflorum only). Adult stage leaves clustered, alternate or whorled, upper surface glossy, green to yellow, lower surface generally paler; stipules absent.  Inflorescences mostly terminal, acrotonic or basitonic, paniculate to single-flowerd. Flowers campanulate, rotate or salverform, aromatic, with parts in 5s regularly placed around the pistil; male inflorescences more floriferous. Sepals mostly spreading and usually less than a third of petal length. Petals ligulate-spathulate, centrally cohering until anthesis or spreading, white-cream becoming yellower with age, rarely red, darkening with age (P. oreillyanum), sometimes bicoloured cream-red (P. bicolor). Stamens free, exserted in male flowers; anthers dorsifixed or basifixed or rarely versatile, always shorter than filaments, ovate to rectangular, dehiscing by apical slits, pollen usually yellow. Staminate flowers can appear perfect with a much-reduced, usually thin, spindle-shaped pistil. Pistil usually with a receptacle disc and basal nectary; ± stipitate; ovary  bi-(or rarely tri-)carpellate but appearing unilocular through the absent septum; placentation parietal, ranging from flat on walls to almost meeting in the centre, ovules inserted in 2 rows per loculus. Pistillate flowers with stigma well developed, prominent; stamens barely reaching to base of style, anthers very thin-sagittate, brownish, nil pollen. Fruit capsular, appearing unilocular, with mainly 2 or occasionally 3 valves in Australia (trivalved in New Zealand species), loculicidally dehiscent or indehiscent (Citriobatus group) or 'tardily' dehiscent; epicarp thin leathery and green-brown, or thick (rarely woody in Australia) and orange. Seeds glossy, red, orange or red-brown, chunky, not winged, packaged in a resinous mass for easy predation, the resin either thin and liquid or thick-coagulating.
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Evergreen, erect, aromatic, shrubs or trees, sometimes epiphytic. Growth in flushes, terminal bud protected by cataphylls (and/or bud scales?). Leaves ex-stipulate, spirally arranged, often crowded in pseudo-whorls, specially towards the ends of the twigs, petioled or seldom sessile, entire (often sinuate in 7. P. sinuatum), generally more or less acuminate, penninerved, young leaves often pubescent. Inflorescences terminal or axillary, in some spp. mainly ramiflorous, few to many-flowered, in fascicles, thyrsoid, or in more or less condensed racemes, rarely solitary, essentially cymose. Flowers bisexual or functionally unisexual apparently to various degree, generally white or pale yellow, except in 5. P. berberidoides, fragrant. Sepals 5, free or connate to various degree. Petals 5, generally ligulate, free or coherent in the lower part, free segments spreading or recurved. Stamens 5, free or occasionally coherent with the corolla tube; filaments in more ♂ flowers slender, in more ♀ flowers somewhat shorter and slightly broadened at the base; anthers dorsifix, introrse, in more ♂ flowers generally oblong, in more ♀ flowers sagittate, and smaller. Pistil sessile or stipitate, stipe thick, 5-furrowed, generally passing into the ellipsoid ovary; ovary glabrous or pubescent, 1-celled, in more ♀ flowers slender or cuneiform, in more ♂ flowers plumper and thicker; placentas 2(-5), (in one case 6!), parietal or basal, with a varying number of ovules; style glabrous, in more ♀ flowers somewhat longer than in more female flowers; stigma in more ♂ flowers hardly thickened, in more female flowers capitate, 2-5-lobed. Capsule globose, ellipsoid or ovoid, 1-celled, loculicidally lengthwise 2(-5)-valved, in Mal. spp. glabrous, mostly orange when mature, often mucronate by the remains of the style; valves of varying thickness, woody or coriaceous, occasionally with resini-ferous cavities, chambers of ducts, inside generally transversely ribbed or pinnately striated; funicles along the median longitudinal placenta or basal. Seeds 1-∞, generally compressed against each other and irregularly angular and wrinkled, variably in size and shape, reddish or blackish, coated by a resinous, viscid fluid. Endosperm horny, large; embryo minute; cotyledons 2-5; germination epigeous.
Trees, shrubs, or sometimes subshrubs, evergreen, pubescent or glabrous. Leavs alternate, appearing opposite or pseudoverticillate, usually clustered at branchlet apex; leaf blade leathery or sometimes membranous, margin entire, undulate-dentate, or rugose. Inflorescences terminal or axillary, umbellate, corymbose, paniculate, or a solitary flower. Flowers bisexual, rarely polygamous. Sepals 5, free, usually short and small. Petals 5, free or partly connate. Stamens 5; filament glabrous; anther dorsifixed, ± sagittate, longitudinally dehiscent. Ovary superior, usually stipitate, of 2 or 3(–5)carpels, 1-loculed or incompletely 2–5-loculed, pubescent or glabrous; ovules usually numerous, sometimes 1–4; placentas parietal and equal in number to carpels, or basilar owing to reduction of ovules. Style short, simple or 2–5-lobed, usually persistent. Capsule ellipsoid or globose, sometimes compressed, dehiscing by 2–5 valves; pericarp woody or leathery, usually with horizontal striae adaxially. Seeds usually surrounded by glutinous or greasy material.
Trees or shrubs, never climbing or spiny. Leaves entire or (outside Africa) rarely undulate or sinuate, glabrous or hairy. Inflorescences usually many-flowered subracemose or subumbellate panicles, terminal and/or axillary from the uppermost leaves, but (outside Africa) flowers sometimes solitary or clustered on the old wood. Flowers regular, up to 15 mm. long, of various colours (but only white, greenish or yellow in Africa), sweet-scented, very often, perhaps always, functionally unisexual; ♂ with long filaments, fertile anthers and slender sterile ovary; ♀ with short filaments, reduced sterile anthers and stout fertile ovary. Ovary unilocular, 2(–5)-merous; style ± as long as ovary, with truncate or sublobed stigma, splitting in fruit according to number of carpels; ovules ovoid. Fruit a capsule; valves leathery or woody, yellow or brown, finally suberect, spreading or reflexed. Seeds variously deformed by mutual pressure, 2–4(–many outside Africa) in two rows on each placenta, ripening orange or red, covered with a sticky slow-drying resin.
Fls perfect to unisexual by abortion, panicled to umbellately arranged or in fascicles, or solitary. Ovary 2-5-celled, style short; ovules few to ∞, sts aborted. Fr. a globose to ovoid or obovoid capsule with solitary to ∞ seeds; valves 2-5, us. woody or leathery, crowned by hardened remains of style; seeds often immersed in viscid fluid. Trees and shrubs with alt. to subverticillate lvs. Genus of some 160 spp., mainly tropical and subtropical southern hemisphere; the N.Z. spp endemic.
Flowers actinomorphic, never more than 15 mm. long, of various colours (in our area only white, yellowish or greenish) sweet-scented, functionally unisexual; male with long filaments, fertile anthers and slender sterile ovary; female with short filaments, reduced sterile anthers and stout fertile ovary.
Ovary 1-locular, with 2–5 (in our area always 2) carpels; style short with a capitate or 2-lobed stigma, splitting in fruit according to the number of carpels; ovules ovoid.
Inflorescences usually variously paniculate or subracemose, terminal or terminal with axillary branches, rarely fascicled, or flowers solitary and axillary.
Seeds variously deformed by mutual pressure, 2-many in 2 rows on each parietal placenta, orange to red, covered with a sticky, slow-drying resin.
Fruit capsular with entire, leathery or woody, erect or spreading valves, sometimes reflexed when quite ripe.
Leaves usually entire, sometimes undulate, rarely subserrate or shallowly lobed, glabrous or pilose.
Petals with claws sometimes connivent and blades usually spreading or revolute.
Usually all the flowers of one inflorescence alike (? monoecious or dioecious).
Sepals free or ± connate, imbricate or not.
Trees or shrubs, never climbing or spiny.
Anthers opening by slits.
Life form -
Growth form
Growth support -
Foliage retention evergreen
Sexuality
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 30.0
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Nitrogen fixer -
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Environment

Malaysian Pittosporums are generally small to at most medium-sized shrubs or trees from the substage of the rain-forest. They do not tend to be social or gregarious but none of them seems to be rare. Only two species are found in forests which are subject to a distinct dry season and that apparently only in part of their area, viz P. moluccanum in the mountain areas of Central and East Java and the Lesser Sunda Islands and P. ferrugineum which is also widely distributed and is the only species occurring in Javanese teak-forests. Both of them may be pioneering in open, grassy mountain forest; P. ferrugineum is, according to VAN DER PIJL ( VAN DER PIJL Ann. Jard. Bot. Btzg 48 1938 136, 138 ), a common pioneer on the lavas of Mt Guntur (W. Java); it has also been employed in re-afforestation work, according to KOORDERS. A similar position is occupied in the Philippines by P. pentandrum which is frequent in parang vegetation.As to altitude many species are rather tolerant; P. sinuatum and P. ramiflorum, for example, range from sea-level through the tropical and montane zones to the lower part of the subalpine zone up to respectively 2700 and 3200 m altitude. There are only two true mountain species, both endemic in New Guinea, viz 6. P. pullifolium (600-)1200-3800 m and 5. P. berberidoides 3000-3900 m. The former species exhibits in its leaves a certain vegetative adaptation towards high altitudes in getting thicker and smaller with recurving leaf margins and condensed habit in proportion to altitude. A similar behaviour is observed in P. ferrugineum, the high-altitude form of which was described as P. versteeghii.Several species have been observed to grow as epiphytes or hemi-epiphytes in tall forest but are also recorded as terrestrial shrubs in more exposed spots.The same species have occasionally been mentioned to be medium-sized trees. There is a possibility that these records are erroneous and that the epiphytic habit was overlooked, the size taken from that of the host tree. Collectors should check this in the field (cf. P. pullifolium)
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Hardiness (USDA) 8-12

Usage

Used in horticultural, including species imported from New Zealand (mainly Pittosporum eugenioides, P. tenuifolium) which are used for hedging and other foliage displays. Pittosporum crassifolium (New Zealand) and P. tobira (widespread in Asia) are particularly useful in heavily polluted areas such as inner city streetscapes. The species known to the Australian horticultural trade as the Queensland Pittosporum (P. rhombifolium) is the type species for genus Auranticarpa (Cayzer et al. 2000b). Pittosporum angustifolium (previously known by the misapplied name P. phillyreoides) is often grown by native plant enthusiasts, while P. undulatum (Sweet Pittosporum) is the most commonly grown of the Australian species but is now a declared weed on three continents including some parts of Australia where it is not native. A few species have a limited use in cabinetry.
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Uses. Several Pittosporums are well known to gardeners because of the modest size of the shrubs or small trees, their attractive-scented flowers and orange fruits. Of the Malaysian species the timber is not available in large sizes or quantities and besides does not seem to be durable. The fruits of some species are here and there employed for medicinal purpose against a multitude of illnesses, e.g. 3. P. resiniferum. Fruits may contain resin and aromatic oil in such quantity that they are inflammable (petroleum nut), for example of 3. P. resiniferum. Leaves and fruits are used for fish poisoning, their bitter substances are obviously due to saponins.
Uses medicinal oil poison timber
Edible -
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Human toxicity -
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Cultivation

Mode -
Germination duration (days) 30 - 60
Germination temperacture (C°) 12
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Images

Pittosporum unspecified picture
Pittosporum unspecified picture

Distribution

Pittosporum world distribution map, present in Argentina, Australia, Bahamas, China, Fiji, Micronesia (Federated States of), Indonesia, Iceland, Japan, Korea (Republic of), Malaysia, New Zealand, Philippines, Papua New Guinea, Tonga, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:326028-2
WFO ID wfo-4000029927
COL ID 6R28
BDTFX ID 100732
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Cylbanida Tobira Pittosporoides Pittosporum Pseuditea Schoutensia Citriobatus

Lower taxons

Pittosporum humile Pittosporum ferrugineum Pittosporum flocculosum Pittosporum halophylum Pittosporum heckelii Pittosporum hematomallum Pittosporum gagnepainianum Pittosporum gatopense Pittosporum malaxanii Pittosporum letocartiorum Pittosporum lancifolium Pittosporum koghiense Pittosporum echinatum Pittosporum eriocarpum Pittosporum crassifolium Pittosporum confertiflorum Pittosporum coriaceum Pittosporum collinum Pittosporum croceum Pittosporum mildbraedii Pittosporum multiflorum Pittosporum muricatum Pittosporum balfourii Pittosporum arborescens Pittosporum aneityense Pittosporum ambrense Pittosporum brevispinum Pittosporum cacondense Pittosporum campbellii Pittosporum bouletii Pittosporum brackenridgei Pittosporum boninense Pittosporum raivavaeense Pittosporum ramiflorum Pittosporum poumense Pittosporum napaulensis Pittosporum naruaiao Pittosporum neelgherrense Pittosporum oblongilimbum Pittosporum perahuense Pittosporum pickeringii Pittosporum parvifolium Pittosporum pancheri Pittosporum paniculatum Pittosporum viridiflorum Pittosporum xanthanthum Pittosporum xenicum Pittosporum sylvaticum Pittosporum suatinum Pittosporum tetraspermum Pittosporum silamense Pittosporum simsonii Pittosporum sinuatum Pittosporum undulatum Pittosporum kaalense Pittosporum lanipetalum Pittosporum inopinatum Pittosporum linearifolium Pittosporum luteum Pittosporum moricrei Pittosporum moluccanum Pittosporum dasycaulon Pittosporum deplanchei Pittosporum michiei Pittosporum baudouinii Pittosporum artense Pittosporum pronyense Pittosporum pullifolium Pittosporum resiniferum Pittosporum ramosii Pittosporum rapense Pittosporum rarotongense Pittosporum rhytidocarpum Pittosporum oreophilum Pittosporum oligodontum Pittosporum obovatum Pittosporum orohenense Pittosporum oubatchense Pittosporum nubicola Pittosporum paniense Pittosporum viridulum Pittosporum spinescens Pittosporum verrucosum Pittosporum sessilifolium Pittosporum trilobum Pittosporum intermedium Pittosporum gracile Pittosporum macrosepalum Pittosporum maireaui Pittosporum bernardii Pittosporum loniceroides Pittosporum leroyanum Pittosporum ornatum Pittosporum spissescens Pittosporum serpentinum Pittosporum anggiense Pittosporum anamallayense Pittosporum aliferum Pittosporum samoense Pittosporum scythophyllum Pittosporum reticosum Pittosporum yunckeri Pittosporum taitense Pittosporum tonkinense Pittosporum takauele Pittosporum tenuivalve Pittosporum tanianum Pittosporum salicifolium Pittosporum polyspermum Pittosporum argentifolium Pittosporum floccosum Pittosporum gayanum Pittosporum glabrum Pittosporum hawaiiense Pittosporum hosmeri Pittosporum kauaiense Pittosporum monae Pittosporum terminalioides Pittosporum goetzei Pittosporum pentandrum Pittosporum divaricatum Pittosporum bracteolatum Pittosporum erioloma Pittosporum fairchildii Pittosporum ellipticum Pittosporum ralphii Pittosporum phillyreoides Pittosporum bicolor Pittosporum oreillyanum Pittosporum umbellatum Pittosporum virgatum Pittosporum trigonocarpum Pittosporum kweichowense Pittosporum leptosepalum Pittosporum daphniphylloides Pittosporum kwangsiense Pittosporum undulatifolium Pittosporum johnstonianum Pittosporum brevicalyx Pittosporum napaulense Pittosporum bullato-ferrugineum Pittosporum humbertii Pittosporum verticillatum Pittosporum stenopetalum Pittosporum pangalanense Pittosporum napaliense Pittosporum patulum Pittosporum turneri Pittosporum obcordatum Pittosporum rigidum Pittosporum crassicaule Pittosporum lineare Pittosporum anomalum Pittosporum cornifolium Pittosporum kirkii Pittosporum revolutum Pittosporum rubiginosum Pittosporum venulosum Pittosporum dallii Pittosporum eugenioides Pittosporum tenuivalvatum Pittosporum paniculiferum Pittosporum balansae Pittosporum kerrii Pittosporum rehderianum Pittosporum xylocarpum Pittosporum subulisepalum Pittosporum pauciflorum Pittosporum omeiense Pittosporum glabratum Pittosporum podocarpum Pittosporum parvicapsulare Pittosporum illicioides Pittosporum elevaticostatum Pittosporum parvilimbum Pittosporum henryi Pittosporum perglabratum Pittosporum planilobum Pittosporum saxicola Pittosporum fulvipilosum Pittosporum kunmingense Pittosporum heterophyllum Pittosporum lenticellatum Pittosporum reflexisepalum Pittosporum merrillianum Pittosporum truncatum Pittosporum leratii Pittosporum qinlingense Pittosporum peridoticola Pittosporum karnatakense Pittosporum rangitahua Pittosporum angustilimbum Pittosporum fulvo-tomentosum Pittosporum macrophyllum Pittosporum viscidum Pittosporum tinifolium Pittosporum abyssinicum Pittosporum ochrosiaefolium Pittosporum pervillei Pittosporum huttonianum Pittosporum pimeleoides Pittosporum crispulum Pittosporum perryanum Pittosporum pachyphyllum Pittosporum buchananii Pittosporum pumilum Pittosporum purpureum Pittosporum poueboense Pittosporum praedictum Pittosporum dzumacense Pittosporum cravenianum Pittosporum ceylanicum Pittosporum berberidoides Pittosporum mackeei Pittosporum longisepalum Pittosporum colensoi Pittosporum fasciculatum Pittosporum angustifolium Pittosporum pulchrum Pittosporum oligophlebium Pittosporum tubiflorum Pittosporum viburnifolium Pittosporum senacia Pittosporum cherrieri Pittosporum coccineum Pittosporum tenuifolium Pittosporum tobira