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Cal regular, deeply saucer-shaped, divided to the middle into 5 broad obtuse lobes; cor bilabiate, the tube wide, the upper lip directed forward, 2-lobed, the lateral lobes of the lower lip also directed forward, the median lobe deflexed; stamens 4; sterile filament dilated at the summit, lying under the upper cor-lip; stigma capitate; fr septicidal; seeds angular, rugose; perennial herbs with opposite, petiolate, serrate or incised lvs and small, dull reddish-brown to greenish fls in large, terminal, paniculiform infls; cal not subtended by conspicuous bracts. 150, mainly temperate Eurasia.
Perennial, biennial or annual herbs or small shrubs with square stems. Lvs usually opposite, entire to pinnatisect. Fls in cymes in the axils of upper lvs or bracts, and sometimes forming terminal racemes or panicles, occasionally solitary. Calyx with 5 equal lobes. Corolla tube short, ± globose; limb with 5 lobes, usually 2-lipped but sometimes almost regular. Fertile stamens 4, included or exserted; staminode usually 1 and scale-like, occasionally 0. Capsule globose to ovoid, septicidal. Seeds numerous, ovoid, rugose, small; wing 0.
