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Annual to perennial herbs; stems trailing or erect; latex 0. Lvs in rosettes and alternate on stems, lyrate-pinnatifid, toothed, or entire. Capitula axillary, solitary, on long peduncles. Involucral bracts free, in several series; outermost row small, narrowed to a linear to spathulate appendage; inner rows progressively longer, oblong, without appendages. Receptacle flat, alveolate, ciliate; scales 0. Outer florets ligulate, ♀. Disc florets ☿ or the inner ♂. Achenes obovoid to obconic, with 2 furrows on the dorsal side. Pappus of 2 rows of oblong scarious scales, the outer very short, the inner longer.
Achenes cylindric-turbinate, ribbed or winged, with the lateral wings folding to form 2 linear chambers on one side, sometimes transversally rugose on the other side, with or without a basal coma of long hairs, otherwise villous or glabrous; pappus c. 2-seriate of delicate hyaline scales, the inner pappus of 5–10 scales equalling the achene in length, the outer pappus smaller.
Disk-florets hermaphrodite, numerous, sometimes central florets functionally male; corollas infundibuliform, regularly 5-lobed; anther bases entire or shortly sagittate; style branches thickened-connate nearly to the apex.
Ray-florets female, sometimes neuter, uniseriate; rays strap-shaped, 3-denticulate; staminodes sometimes present; style branches free and lanceolate.
Phyllaries free, many-seriate, scarious-hyaline on the margins, the outer phyllaries usually with apical appendages.
Perennial or annual herbs, araneose-lanate or glandular, often acaulescent.
Receptacle flat, fimbrilliferous-alveolate with walls produced into setae.
Capitula heterogamous radiate, solitary, large and many-flowered.
Leaves alternate or radical, entire to lyrate-pinnatifid.
