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Styles 2 (3–4 have been rarely reported), connivent for most of their length and often with very short divergent stigmatic surfaces at their apex.
Leaves conspicuously petiolate, simple and palmate to digitately compound, often grouped in an “umbrella ” arrangement at the ends of branches.
Fruit subglobose, urceolate, obovoid or even obconical to wedge-shaped (owing to congestion in densely packed spikes), sometimes fleshy.
Trees, shrubs (rarely suffrutices), glabrous to puberulous (more rarely densely pubescent, especially in immature states).
Disk flat, depressed or sometimes fused with the styles to form a well-developed conical Stylopodium.
Leaflets very variable in shape and marginal definition; apices often long-acuminate.
Calyx forming a shallow cup, sometimes with a 4–5-dentate margin.
Seed ovoid or somewhat compressed; endosperm ruminate.
Flowers 4–8 mm. in diameter with (4)5 greenish petals.
Ovary 2-locular (rarely with supernumerary carpels).
Inflorescence usually spicate but may be racemose.
Stipules often quite conspicuous.
Floral bracts small or obsolete.
Stamens (4)5.
