Shrubs or trees, sometimes scrambling. Leaves alternate, sometimes fasciculate on short-shoots, simple. Capitula in dense heads of heads, the glomerules, these solitary or grouped in cymes, each glomerule formed of many capitula and usually subtended by bracts, capitula disciform. Phyllaries 3 per capitulum (primary head); receptacle paleate. Florets 2–6, 2 filiform female and 2–4 functionally male disc florets. Anthers caudate. Achenes 3-angled, often with twin-hairs (hairs of two parallel rows of cells); pappus of scales or absent.
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