Goodeniaceae R.Br.

Family

Angiosperms > Asterales

Characteristics

Mostly perennial or annual herbs, but sometimes subshrubs, shrubs or rarely scramblers, glabrous or hairy with simple or branched or glandular hairs, sometimes viscid; roots usually primary, sometimes adventitious. Leaves mostly alternate or radical, rarely opposite; lamina simple, margin entire to dentate or rarely lobed; stipules absent but often with an axillary tuft of hairs. Inflorescence a dichasium, thyrse, raceme, umbel, spike or a single terminal head surrounded by an involucre of green bracts, or flowers solitary. Flowers zygomorphic or rarely actinomorphic (Brunonia), bisexual, protandrous. Sepals usually 5, rarely 3, sometimes reduced to rim or obsolete, free or sometimes connate, sometimes persistent (Brunonia). Corolla of 5 petals, connate at least to some degree, usually deeply slit (except Brunonia), fanlike or two-lipped, or sometimes pouched, the lobes ± winged, often with auricles on margin of the adaxial slit, the auricles surrounding the indusium. Stamens 5, opposite sepals, epigynous or epipetalous or rarely apparently hypogynous; anthers free or connate into a tube around the style, tetrasporangiate, dithecal, introrse, dehiscing by longitudinal slits. Intrastaminal nectariferous glands sometimes present, glands 1 or 2. Gynoecium, on anatomical evidence, suggested to be of 4 connate carpels. Ovary usually inferior, or sometimes half-inferior or superior (Brunonia, Goodenia), usually 2-locular (septum usually incomplete), rarely 4 or apparently 1; style usually entire with a hollow pollen-cup (indusium) usually enclosing the stigma, often curved; indusium often equipped with hairs or glabrous; stigma 2-fid or unlobed, rarely 2–4-fid. Ovule 1–numerous per locule; placentation axile or basal. Fruit usually a capsule, sometimes a drupe or nut (sometimes with persistent calyx, in Brunonia), rarely separating transversely into 1-seeded woody segments. Seeds 1–numerous, often flattened and winged, with or without endosperm (Brunonia), rarely with a caruncle; testa membranous to thickened; embryo straight.
More
Herbs, shrubs, or rarely small trees [or vines], without laticifers. Stipules absent. Leaves alternate, rarely opposite or whorled, simple. Inflorescences a cyme, raceme, head, or sometimes single flowered in axils. Flowers perfect, protandrous. Calyx epigynous, tubular, mostly (3-)5-lobed. Corolla sympetalous, irregular, 2-lipped or sometimes 1-labiate due to 2-segmentation of posterior lip to base; lobes 5, valvate, often induplicate. Stamens 5, alternate with corolla lobes, free from corolla or adnate to base of corolla tube; anthers 4-sporangiate and dithecal, introrse, longitudinally dehiscent. Ovary inferior, semi-inferior [or rarely (Velleia Smith) essentially superior], (1 or)2-locular [occasionally 4-locular (Scaevola porocarya F. Mueller)]; ovules 1 to many per locule, on axile or basal-axile placentas, anatropous. Fruit usually a capsule, less often a drupe or a small nut, often with persistent calyx. Seeds with a straight embryo embedded in copious endosperm.
Trees, shrubs or herbs, without latex. Leaves alternate, rarely opposite, or radical, exstipulate, simple, often obovate to spathulate. Inflorescence cymose or reduced to solitary axillary flowers. Flowers hermaphrodite, protandrous, irregular. Calyx adnate to the ovary basally, distally free as a 5-toothed or entire limb. Corolla with a long tube split for part or all of its length and a limb divided into 5 subequal lobes. Stamens 5, from the base of the corolla, free or occasionally joined at the anthers; anthers dehiscing introrsely. Ovary superior to inferior, 1–2-locular, each containing 1 or several ovules; style emerging laterally from the split corolla-tube, bearing a cup-shaped ciliate indusium around the simple or bifid stigma. Fruit a capsule or drupe. Seeds with endosperm
Stamens 5, alternating with the corolla–lobes and inserted at the base of the corolla, usually free but sometimes joined with the anthers; anthers 2–thecous, with the theca parallel and opening longitudinally
Stamens 5, alternate with the corolla-lobes, free or rarely shortly adnate to the corolla; anthers 2-celled, free or connivent around the style
Style simple or rarely divided, emerging laterally from the split corolla–tube; stigma truncate or bilobed, with an apical indusium
Flowers sometimes solitary and axillary but more often arranged in a spike, raceme or panicle, hermaphrodite, protandrous
Calyx tubular, adnate to the ovary or very rarely free; limb consisting of 5 persistent lobes, rarely entire
Corolla sympetalous, 5–merous, bilabiate or very rarely 1–lipped; tube long, often villous inside
Leaves usually alternate, rarely opposite, sometimes all radical, entire, simple, exstipulate
Corolla gamopetalous, bilabiate or rarely 1-lipped, lobes valvate, often induplicate
Herbs, small shrubs or rarely shrubs or small trees without latex, rarely spinescent
Seeds with endosperm, small, flat, with a thin or thick and often crustaceous testa
Ovary mostly inferior, 1–2 (–4)–locular; ovules 1–several, erect or ascending
Seeds small, with straight embryo in the middle of copious endosperm
Ovary mostly inferior, 1-4-celled; stigma indusiate at the top
Leaves alternate or rarely opposite; stipules absent
Ovules 1 or more in each cell, erect or ascending
Calyx tubular, adnate to the ovary, rarely free
Fruit drupaceous or nut-like, or capsular
Fruit a drupe, capsule or nutlike
Lobes valvate, subequal
Disc sometimes present
Herbs or undershrubs
Flowers zygomorphic
Life form
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Sexuality hermaphrodite
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Root system adventitious-root
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Environment

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Hardiness (USDA) 7-11

Usage

Goodenia, Lechenaultia and Scaevola are grown as ornamental plants, e.g. cultivar Fan-flowers commonly used as groundcovers. Goodenia ovata (Hop Goodenia) is often used in revegetation projects in urban areas in southeastern Australia. Several species have medicinal uses. An extract from Goodenia ovata was used by Aboriginal people as a soporific for their children (HortFlora). Scaevola taccada (Beach Naupaka) is coastal shrub sometimes used to prevent beach erosion (Christenhusz et al. 2017).
Uses medicinal ornamental
Edible -
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Cultivation

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Images

Goodeniaceae unspecified picture
Goodeniaceae unspecified picture

Distribution

Goodeniaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:77126671-1
WFO ID wfo-7000000255
COL ID 623M9
BDTFX ID -
INPN ID 445293
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Goodeniaceae

Lower taxons

Scaevola Anthotium Crossotoma Leschenaultia Pentaptilon Diaspasis Goodenia Leschenaultia Selliera Verreauxia Velleia Brunonia Coopernookia Dampiera