Mericarps indehiscent, 1-seeded, hemispheric-reniform, conical, truncate or compressed-orbicular, winged or wingless, often auricled at base, the outer face reticulate, rugose, spinescent or more or less smooth.
Leaves alternate, opposite or fasciculate, linear to orbicular, entire, sessile or petiolate, sometimes subsucculent, exstipulate.
Inflorescences terminal or lateral, cymose, often umbelliform, sessile or pedunculate, few-or many-flowered, lax or dense.
Stamens (5)–7, hypogynous; filaments broadened below, inserted usually on hypogynous disk.
Staminodes 0–5, generally petaloid, arising from the base of the stamens, free.
Glabrous or viscid, glandular-pubescent annual or perennial herbs or subshrubs.
Flowers hermaphrodite, whitish or greenish, small, more or less pedicellate.
Ovary superior, syncarpous, bicarpellate, 2-locular; style 1; stigmas 2.
Perianth-segments 5, generally unequal, herbaceous, free.
Fruit separating into 2 mericarps.