Leaves very rarely acuminate, with a more or less rounded extra-floral nectary on the upper surface at the base of the midrib, and sometimes at the base of the lower lateral nerves; midrib and lateral nerves depressed above and prominent beneath; reticulation very dense, isodiametrical, scarcely to strongly prominent beneath, in the latter case turning the interreticular areoles into deep cavities; indumentum extremely variable; hairs simple, penicillate, or stellate, straight, curved, curled, or coiled, from very short to long, with punctiform spherical glands, sometimes making the lamina viscid, abundant or dispersed on the two surfaces of the leaf.
Fruit subglobose, with a hard thick pericarp and with 1 cavity containing 1 seed, surrounded by the 5 subequal wings of the accrescent calyx; wings with minute penicillate hairs outside and inside.
Flowers small, in axillary clusters, with very hairy sepals and petals; receptacle produced into a very short broad androgynophore.
Stamens numerous; anthers sohrt, very often produced at the apex into a triangular or ovate appendage.
Ovary hairy, completely 3-locular with 2 ovules in each loculus.
Shurbs to medium-sized trees, without buttresses.
Calyx-lobes equally enlarged in fruit.
Anthers with produced connective
Stamens numerous
Petals 5, hairy
Calyx-lobes 5