Corolla small, often distinctly apiculate in bud (due to lobe appendages); usually yellow or greenish-yellow; corolla tube shortly cylindrical or campanulate, glabrous or hairy outside, with a ring of deflexed hairs inside and densely hairy at the throat; lobes 5, narrowly triangular, mostly reflexed, sometimes each with a distinct narrow appendage at the apex.
Inflorescences usually many-flowered, in axillary, divaricately branched cymes at opposite sides of nodes, often at leafless nodes and sometimes flowering before leaves are fully developed; cyme branches ± elongate with regularly spaced shortly petiolate flowers.
Leaves paired, never whorled, often quite large; stipules broad and connate into a short sheath at the base, produced into a linear or subulate appendage at the apex, hairy inside, deciduous or more or less persistent.
Fruit large, indehiscent and fleshy, ± globose, often lobed or angular in the dry state, (4)5(6)-locular, containing (4)5(6) pyrenes, sometimes bearing the remains of calyx lobes, glabrous, often edible.
Calyx tube hemispherical or depressed campanulate; calyx limb divided into 5(6) triangular, oblong, linear or ligulate lobes.
Style filiform-cylindrical, glabrous, shortly exserted; pollen presenter cylindrical, very shortly 5-lobed at the apex.
Deciduous shrubs or small trees, unarmed or rarely spiny, with glabrous to densely velvety foliage.
Ovary 5-locular; ovule solitary in each locule, pendulous.
Anthers ovate or oblong, apiculate, shortly exserted.
Disk annular, slightly raised.