Scutellaria L.

Skullcaps (en), Scutellaires (fr), Scutellaire (fr)

Genus

Angiosperms > Lamiales > Lamiaceae

Characteristics

Herbs or subshrubs, rarely shrubs, not aromatic. Leaves entire to pinnatifid. Inflorescences terminal or axillary racemes or spikes; floral leaves usually bractlike apically. Flowers axillary, opposite or sometimes alternate apically. Calyx short tubular, dorsiventrally flattened, 2-lipped; lips entire, closed and ultimately divided to base along sutures in fruit; upper lip deciduous, with a transverse, rounded, concave, scalelike scutellum (shield) or without and abaxially conspicuously saccate; lower lip persistent. Corolla 2-lipped; tube exserted, arcuate or suberect, gradually widening to throat, base bent and saccate or spurred, usually puberulent annulate outside; upper lip erect, galeate; lower lip 3-lobed, middle lobe broad, flattened, entire, lateral lobes ± joined to upper lip and sometimes spreading. Stamens 4, didynamous, anterior 2 longest, underlying upper lip; anthers close together in pairs, bearded on cell aperture; posterior pair conspicuously 2-celled, ± apically acute, anterior pair 1-celled by abortion. Style subulate, apically unequally 2-cleft. Nutlets oblate, globose, to ovoid.
More
lip deciduous and usually with a transverse squama becoming an erect scutellum at maturity, the lower lip persistent; corolla tubular, bilabiate, generally long-exserted, red, orange, yellow, blue, violet, or white and one of the previous colors, the tube cylindrical and ? sigmoid, arcuate, or more sharply bent, the upper lip cup-shaped or galeate and ? joined obliquely to the lateral lobes of the lower lip, glabrous or pubescent within or without; stamens 4, paired, included in the upper lip, the pairs attached at different levels on the corolla tube, the filaments distinct, the connective not developed, the lower anthers with 1 functional theca; ovary 4-lobed, gynobase generally conspicuous and asymmetrical, variously de-scribed or functioning as a gynophore, the style long and slender, bifid near the tip, the upper style branch small or not apparent. Nutlets ca 3-4, attached at the base, + ovoid or spherical, the pericarp variously tubercular or verrucose, some-times provided with a band near the equator.
Herbs, sometimes woody below. Leaves opposite, petioled, rarely sessile. Flowers in terminal or upper axillary racemose inflorescence. Bracts minute or conspicuous and foliaceous. Calyx short, campanulate, 2-lipped, accrescent; calyx tube with a large caducous shield-or pouch-like appendage ('scutellum') above the upper lip; in fruit, at first the lips closed together, then the upper lip falling away together with the appendage. Corolla trumpet-shaped, usually sharply recurved from the base and erect upwards, not annulate within, 2-lipped; upper lip often boat-shaped, entire or notched; lower lip broad, 3-lobed. Stamens 4; lower pair longer, anthers often dimidiate or 1-celled; upper pair shorter, anthers 2-celled. Disk tubular, elongate. Ovary oblique, on a short gynophore; style 2-fid. Nutlets very minute, smooth, granular or hispid; seeds more or less transverse, with curved embryo.
Cal obscurely nerved, strongly bilabiate, with entire lips, bearing a prominent transverse appendage on or proximal to the upper lip; cor 2-lipped, the upper lip generally ± galeate, the lateral lobes of the lower lip somewhat separated from the broad central lobe and marginally partly connate with the upper lip; stamens 4, ascending under the upper lip, the anthers of the upper pair with 2 pollen sacs, those of the lower pair with only one; style gynobasic, its upper branch reduced or wanting; ours all perennial, mostly rhizomatous; fls blue or violet, seldom pink or white, solitary in the axils or in terminal or axillary racemes. 300, cosmop.
Usually herbs, sometimes small shrubs, not aromatic, sometimes rhizomatous. Lvs simple, entire to pinnatisect. Fls axillary, one to each opposite lf in upper axils or in terminal spikes or racemes. Bracts large and leaflike or reduced. Calyx 2-lipped; tube campanulate, not gibbous at base; upper lip with broad scale or scutellum which is eventually deciduous; lower lip persistent. Corolla tube usually > limb, often curved upwards, glabrous or ± hairy inside; limb 2-lipped. Stamens didynamous and parallel, lying beneath upper lip. Style gynobasic, its branches unequal. Nutlets subglobose or flattened, often granular or tuberculate.
Fls solitary or in spikes or racemes. Calyx with entire lips, upper with dorsal scale. Corolla-tube elongate, dilated towards 2-lipped limb; upper lip entire, lower 3-lobed. Anthers ciliolate; lower 1-celled, upper 2-celled. Nutlets small. Annual to perennial herbs or subshrubs. A widespread genus of some 250 spp., the N.Z. sp. endemic.
Herbs, mostly perennials, or half shrubs, erect or procumbent, rarely climbers, glabrous or pubescent. Leaves simple, opposite, petiolate or subsessile, pinnately-
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

In the Malesian tropics both at low and high altitude.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 5-12

Usage

Uses medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 14 - 180
Germination temperacture (C°) 10
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Scutellaria unspecified picture

Distribution

Scutellaria world distribution map, present in Afghanistan, Åland Islands, Albania, Andorra, Austria, Burkina Faso, Bangladesh, Bulgaria, Bahamas, Belarus, Belize, Bolivia (Plurinational State of), Central African Republic, Switzerland, Chile, China, Colombia, Costa Rica, Cuba, Cyprus, Czech Republic, Germany, Djibouti, Dominica, Denmark, Algeria, Ecuador, Eritrea, Spain, Ethiopia, Finland, Falkland Islands (Malvinas), France, Micronesia (Federated States of), Georgia, Ghana, Guinea, Greece, Guatemala, Honduras, Haiti, Hungary, Indonesia, Ireland, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Japan, Kazakhstan, Kenya, Kyrgyzstan, Lao People's Democratic Republic, Libya, Morocco, Moldova (Republic of), Mali, Myanmar, Montenegro, Mozambique, Malaysia, Nigeria, Nicaragua, Netherlands, Norway, Nepal, New Zealand, Pakistan, Panama, Peru, Philippines, Palau, Poland, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Paraguay, Romania, Russian Federation, Rwanda, Saudi Arabia, Sudan, Senegal, Singapore, Sierra Leone, Somalia, Sweden, Turks and Caicos Islands, Chad, Thailand, Turkmenistan, Tunisia, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Ukraine, Uruguay, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, Yemen, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:30003498-2
WFO ID wfo-4000034856
COL ID 8W3H6
BDTFX ID 87191
INPN ID 197540
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Scutellaria Perilomia Hastifolia

Lower taxons

Scutellaria adenostegia Scutellaria agrestis Scutellaria alborosea Scutellaria altamaha Scutellaria amicorum Scutellaria amoena Scutellaria angustifolia Scutellaria anitae Scutellaria anomala Scutellaria antirrhinoides Scutellaria arabica Scutellaria aramberrana Scutellaria arenicola Scutellaria argentata Scutellaria arguta Scutellaria ariana Scutellaria artvinensis Scutellaria aurea Scutellaria axilliflora Scutellaria baicalensis Scutellaria barbata Scutellaria bolanderi Scutellaria bornmuelleri Scutellaria botschantzevii Scutellaria brachyspica Scutellaria brittonii Scutellaria bushii Scutellaria californica Scutellaria cardiophylla Scutellaria caryopteroides Scutellaria catharinae Scutellaria caucasica Scutellaria caudifolia Scutellaria chihshuiensis Scutellaria chimenensis Scutellaria chungtienensis Scutellaria x churchilliana Scutellaria coccinea Scutellaria cochinchinensis Scutellaria cordifolia Scutellaria daghestanica Scutellaria discolor Scutellaria drummondii Scutellaria edelbergii Scutellaria elliptica Scutellaria filicaulis Scutellaria flabellulata Scutellaria gardoquioides Scutellaria gaumeri Scutellaria glabriuscula Scutellaria guilielmi Scutellaria guttata Scutellaria haesitabunda Scutellaria hainanensis Scutellaria havanensis Scutellaria hintoniorum Scutellaria hissarica Scutellaria holosericea Scutellaria honanensis Scutellaria immaculata Scutellaria incarnata Scutellaria indica Scutellaria inflata Scutellaria integrifolia Scutellaria intermedia Scutellaria isocheila Scutellaria juzepczukii Scutellaria kamelinii Scutellaria karatavica Scutellaria karkaralensis Scutellaria kotkaiensis Scutellaria kuromidakensis Scutellaria kurssanovii Scutellaria laeteviolacea Scutellaria laevis Scutellaria leptosiphon Scutellaria leptostegia Scutellaria leucantha Scutellaria lilungensis Scutellaria linarioides Scutellaria linczewskii Scutellaria lindbergii Scutellaria lindeniana Scutellaria linearis Scutellaria lipskyi Scutellaria litwinowii Scutellaria longifolia Scutellaria longituba Scutellaria lotienensis Scutellaria lundellii Scutellaria lutea Scutellaria luteocaerulea Scutellaria lutescens Scutellaria macrochlamys Scutellaria maxonii Scutellaria meehanioides Scutellaria megalaspis Scutellaria megalodonta Scutellaria megaphylla Scutellaria mellichampii Scutellaria mesostegia Scutellaria mexicana Scutellaria microdasys Scutellaria microphylla Scutellaria microviolacea Scutellaria mociniana Scutellaria modesta Scutellaria mollifolia Scutellaria mollis Scutellaria montana Scutellaria multicaulis Scutellaria multiflora Scutellaria multiglandulosa Scutellaria muramatsui Scutellaria muriculata Scutellaria nana Scutellaria navicularis Scutellaria nepetifolia Scutellaria nepetoides Scutellaria nervosa Scutellaria neubaueri Scutellaria x neumannii Scutellaria nevskii Scutellaria x nicholsonii Scutellaria nuristanica Scutellaria obtusifolia Scutellaria ochotensis Scutellaria ocmulgee Scutellaria ocymoides Scutellaria oligodonta Scutellaria omeiensis Scutellaria orbicularis Scutellaria orichalcea Scutellaria orizabensis Scutellaria orthocalyx Scutellaria orthotricha Scutellaria ossethica Scutellaria pallidiflora Scutellaria parvula Scutellaria petiolata Scutellaria physocalyx Scutellaria pingbienensis Scutellaria pinnatifida Scutellaria poecilantha Scutellaria poliochlora Scutellaria polyadenia Scutellaria pontica Scutellaria popovii Scutellaria pseudoserrata Scutellaria pseudotenax Scutellaria purpurascens Scutellaria purpureocardia Scutellaria pycnoclada Scutellaria quadrilobulata Scutellaria repens Scutellaria rosei Scutellaria roseocyanea Scutellaria rubromaculata Scutellaria rubropunctata Scutellaria russelioides Scutellaria salviifolia Scutellaria sapphirina Scutellaria saxatilis Scutellaria scandens Scutellaria schachristanica Scutellaria schugnanica Scutellaria schweinfurthii Scutellaria sciaphila Scutellaria scutellarioides Scutellaria semicircularis Scutellaria sessilifolia Scutellaria shansiensis Scutellaria shikokiana Scutellaria shweliensis Scutellaria sibthorpii Scutellaria sieberi Scutellaria sieversii Scutellaria siphocampyloides Scutellaria stachyoides Scutellaria stenosiphon Scutellaria strigillosa Scutellaria tarokoensis Scutellaria tatianae Scutellaria tayloriana Scutellaria tenax Scutellaria tenera Scutellaria tenuiflora Scutellaria ternejica Scutellaria texana Scutellaria theobromina Scutellaria tienchuanensis Scutellaria titovii Scutellaria toguztoravensis Scutellaria tomentosa Scutellaria tournefortii Scutellaria tsinyunensis Scutellaria tuberifera Scutellaria tuberosa Scutellaria tubiflora Scutellaria tuminensis Scutellaria turgaica Scutellaria tuvensis Scutellaria uliginosa Scutellaria velutina Scutellaria villosissima Scutellaria violacea Scutellaria vitifolia Scutellaria volubilis Scutellaria weishanensis Scutellaria wendtii Scutellaria wenshanensis Scutellaria wightiana Scutellaria wongkei Scutellaria wrightii Scutellaria xanthosiphon Scutellaria xylorrhiza Scutellaria zaprjagaevii Scutellaria zivari Scutellaria viscidula Scutellaria jaliscana Scutellaria adenotricha Scutellaria altaica Scutellaria andamanica Scutellaria andrachnoides Scutellaria androssovii Scutellaria angrenica Scutellaria bambusetorum Scutellaria chaematochlora Scutellaria chamaedrifolia Scutellaria chekiangensis Scutellaria chenopodiifolia Scutellaria chevalieri Scutellaria cypria Scutellaria delavayi Scutellaria dependens Scutellaria floridana Scutellaria fontqueri Scutellaria forrestii Scutellaria fraxinea Scutellaria galericulata Scutellaria glabra Scutellaria glandulosa Scutellaria grossa Scutellaria grossecrenata Scutellaria guatemalensis Scutellaria heterophylla Scutellaria heterotricha Scutellaria heydei Scutellaria hintoniana Scutellaria humilis Scutellaria hunanensis Scutellaria hypericifolia Scutellaria khasiana Scutellaria kikai-insularis Scutellaria kingiana Scutellaria kiusiana Scutellaria knorringiae Scutellaria lacei Scutellaria lanipes Scutellaria laxa Scutellaria macra Scutellaria moniliorhiza Scutellaria mulleri Scutellaria nigrocardia Scutellaria novae-zelandiae Scutellaria potosina Scutellaria pseudocoleus Scutellaria ramosissima Scutellaria ramazanica Scutellaria regeliana Scutellaria seleriana Scutellaria sosnowskyi Scutellaria spectabilis Scutellaria squarrosa Scutellaria stewartii Scutellaria stocksii Scutellaria striatella Scutellaria subcaespitosa Scutellaria supina Scutellaria taiwanensis Scutellaria tenuipetiolata Scutellaria umbratilis Scutellaria urticifolia Scutellaria utriculata Scutellaria yingtakensis Scutellaria yunnanensis Scutellaria albida Scutellaria likiangensis Scutellaria rehderiana Scutellaria splendens Scutellaria lutilabia Scutellaria microphysa Scutellaria molanguitensis Scutellaria flocculosa Scutellaria franchetiana Scutellaria novorossica Scutellaria oaxacana Scutellaria oreophila Scutellaria phyllostachya Scutellaria formosa Scutellaria fragillima Scutellaria fruticetorum Scutellaria galerita Scutellaria ghorana Scutellaria mongolica Scutellaria monterreyana Scutellaria muzquiziana Scutellaria oschtenica Scutellaria glabrata Scutellaria grandiflora Scutellaria grossheimiana Scutellaria helenae Scutellaria hispidula Scutellaria karjaginii Scutellaria oblonga Scutellaria glaphymstachys Scutellaria glechomoides Scutellaria glutinosa Scutellaria cristata Scutellaria dumetorum Scutellaria persica Scutellaria picta Scutellaria platensis Scutellaria platystegia Scutellaria platystoma Scutellaria polyadena Scutellaria polyphylla Scutellaria polytricha Scutellaria prilipkoana Scutellaria pseudocoerulea Scutellaria raddeana Scutellaria aurata Scutellaria benthamiana Scutellaria comosa Scutellaria darriensis Scutellaria darvasica Scutellaria durangensis Scutellaria ebracteata Scutellaria eplingii Scutellaria farsistanica Scutellaria fauriei Scutellaria resinosa Scutellaria reticulata Scutellaria robusta Scutellaria asperiflora Scutellaria assamica Scutellaria atrocyanea Scutellaria baldshuanica Scutellaria bartlettii Scutellaria blepharophylla Scutellaria bucharica Scutellaria carmenensis Scutellaria chiangii Scutellaria chodshakasiani Scutellaria cyanocheila Scutellaria amphichlora Scutellaria cylindriflora Scutellaria amabilis Scutellaria taipeiensis Scutellaria petersoniae Scutellaria parrae Scutellaria botbaevae Scutellaria slametensis Scutellaria x ketenoglui Scutellaria tortumensis Scutellaria molinarum Scutellaria multiramosa Scutellaria oblongifolia Scutellaria somalensis Scutellaria talamancana Scutellaria anatolica Scutellaria stachys Scutellaria discolor Scutellaria balearica Scutellaria sporadum Scutellaria brevibracteata Scutellaria yildirimlii Scutellaria rubicunda Scutellaria hsiehii Scutellaria cuevasiana Scutellaria sublitoralis Scutellaria tenasserimensis Scutellaria khaoyaiensis Scutellaria dombeyi Scutellaria tutensis Scutellaria chalicophila Scutellaria colebrookeana Scutellaria colpodea Scutellaria cordifrons Scutellaria diffusa Scutellaria gontscharovii Scutellaria goulimyi Scutellaria hirta Scutellaria nigricans Scutellaria nummulariifolia Scutellaria oligophlebia Scutellaria oxystegia Scutellaria yangbiensis Scutellaria austrotaiwanensis Scutellaria violascens Scutellaria indica Scutellaria javanica Scutellaria langbianensis Scutellaria x minkwitziae Scutellaria przewalskii Scutellaria anhweiensis Scutellaria mairei Scutellaria ocellata Scutellaria pacifica Scutellaria rupestris Scutellaria suffrutescens Scutellaria valdiviana Scutellaria pekinensis Scutellaria scordiifolia Scutellaria teniana Scutellaria yezoensis Scutellaria fedtschenkoi Scutellaria sarmentosa Scutellaria araratica Scutellaria araxensis Scutellaria granulosa Scutellaria rhomboidalis Scutellaria sedelmeyerae Scutellaria sevanensis Scutellaria paulsenii Scutellaria albituba Scutellaria attenuifolia Scutellaria aurantiaca Scutellaria lactea Scutellaria saslayensis Scutellaria tucurriquensis Scutellaria woodii Scutellaria formosana Scutellaria playfairii Scutellaria porphyrantha Scutellaria prostrata Scutellaria sichourensis Scutellaria subcordata Scutellaria subintegra Scutellaria swatensis Scutellaria szovitziana Scutellaria talassica Scutellaria tapintzensis Scutellaria tauricola Scutellaria pamirica Scutellaria paradoxa Scutellaria calcarata Scutellaria hookeri Scutellaria incisa Scutellaria incurva Scutellaria inghokensis Scutellaria insignis Scutellaria irrasa Scutellaria iskanderi Scutellaria iyoensis Scutellaria jodudiana Scutellaria macrodonta Scutellaria macrosiphon Scutellaria adsurgens Scutellaria alta Scutellaria krasevii Scutellaria kugarti Scutellaria serboana Scutellaria sipilensis Scutellaria hastifolia Scutellaria serrata Scutellaria lateriflora Scutellaria minor Scutellaria orientalis Scutellaria incana Scutellaria alabamensis Scutellaria costaricana Scutellaria ovata Scutellaria racemosa Scutellaria altissima Scutellaria columnae Scutellaria alpina