Calamus L.

Calamus (en)

Genus

Angiosperms > Arecales > Arecaceae

Characteristics

Solitary or clustering, acaulescent to high-climbing pleonanthic dioecious palms. Stems very slender, only a few mm. in diameter to robust (15 cm. or more), in Africa moderate, 1–3 cm. in diameter, branching sympodially at the base. Leaf-sheaths tightly enclosing the stem, variously armed with spines and spiculae or unarmed, often covered with indumentum; often continued into an ocrea of varying length (in Africa conspicuous). Mature leaf of two kinds, either terminating in a long barbed whip (cirrus) or without such a whip (African and some Asiatic species), species without a cirrus normally but not always bearing a similar barbed whip (flagellum), adnate to the leaf-sheath at the base, equivalent to a modified sterile inflorescence.. Petiole prominent or absent, variously armed with spines and hooks.. Rachis usually armed with reflexed hooks.. Leaflets narrow to broad or rhomboid (praemorse in one Australian species), single-fold, arranged regularly or irregularly on either side of the rachis or variously clustered, fanned or paired, variously hairy, scaly or spiny. Inflorescence axillary, with the base of the peduncle adnate to the internode and sheath of the following leaf, hence appearing in a non-axillary position, branching to 2–4 orders, with or without a long terminal flagellum.. Bracts variously armed, tubular, tightly sheathing, very rarely splitting, sometimes with an expanded limb, never caducous, though rarely tattering and decaying before fruiting; prophyll usually 2-keeled and empty; other bracts on main axis subtending close to very distant partial inflorescences; partial inflorescences bearing bracts subtending rachillae; rachillae usually with approximate tubular bracts, each, except for the one or more basalmost, subtending a flower or flower group.. In ♂ inflorescence flowers solitary, borne together with a bracteole (‘involucre’). In ♀ inflorescence flowers borne in pairs, a sterile ♂ (acolyte) together with a fertile ♀ and 2 bracteoles (‘involucrophore’ and ‘involucre’). Male flower symmetrical; calyx tubular, 3-lobed; corolla 3-lobed, divisions almost reaching the base; stamens 6, epipetalous, with free filaments; pistillode minute or absent. Sterile ♂ flower as the fertile ♂ but anthers empty.. Female flower with calyx and corolla ± as in the ♂ flower; staminodes 6; ovary covered in vertical rows of reflexed scales and tipped with 3 stigmas; locules 3, incomplete, each with a single ovule, normally only 1 ovule developing. Fruit variously shaped, tipped with the stigmatic remains, bearing persistent calyx and corolla basally, and covered in vertical rows of reflexed scales. Seed with thick or thin, sweet or sour or very astringent sarcotesta and variously shaped hard diaspore; endosperm homogeneous or ruminate; embryo basal or lateral.
More
Stems clustered or less often solitary, mostly slender and climbing, occasionally non-climbing and then either short and subterranean or free standing. Leaves 10-60, pinnate, spiny; sheaths closed in climbing stems, open in non-climbers, variously hairy and spiny, hairs of young sheaths soon wearing off; sheath spines scattered or densely arranged (rarely absent), or arranged in rows, occasionally hairy on margins, variously shaped and colored, sometimes arranged in overlapping, interlocking rings and forming chambers; sheath apices usually extended above point of insertion of petiole into an inconspicuous or prominent ocrea; knees present in climbing stems; flagella present in climbers, whiplike and armed with small, grapnel-like spines; cirri present in few species (and then flagella absent, and vice versa); pinnae variously arranged and shaped, occasionally whitish or gray on abaxial surfaces. Plants dioecious. Inflorescences usually elongate, branched to 3 orders, male inflorescences more branched than female ones; branches and rachillae covered with overlapping bracts with clawed spines on outer surfaces in climbers; male flowers borne along opposite sides of rachillae; female flowers borne in pairs with a sterile male flower, along opposite sides of rachillae. Fruits mostly small, variously shaped and colored, usually 1-seeded, sometimes pedicellate, always covered with overlapping scales, these sometimes with a groove down middle; endosperm homogeneous or ruminate; germination adjacent; eophylls bifid or pinnate.
Solitary or clustering, pleonanthic, dioecious, spiny palms, mostly climbing but also erect or acaulescent (not in Australia). Stems slender; internodes short to long. Leaves paripinnate, occasionally terminating in a cirrus; leaf sheath tubular, usually armed, sometimes bearing a flagellum; petiole absent to long, usually armed; rachis usually armed. Pinnae evenly distributed or clustered, opposite or subopposite, variously armed; apices entire, rarely praemorse. Staminate and pistillate inflorescences superficially similar, axillary, basally adnate to adjacently distal internode and sheath of following leaf, compoundly branched, frequently flagellate at tip; bracts tubular, usually armed. Staminate flower solitary; stamens 6, epipetalous; pistillode small or lacking. Pistillate flower borne together with sterile staminate flower to form a dyad; staminodes 6; stigmas 3. Fruit ellipsoid-ovoid to globose, covered in vertical rows of imbricate scales. Seed subglobose, sarcotestal; embryo basal or lateral (not in Australia). Eophyll bifid or paripinnate (not in Australia).
Life form perennial
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Foliage retention deciduous
Sexuality dioecy
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Environment

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Hardiness (USDA) 9-12

Usage

Uses -
Edible -
Therapeutic use -
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Cultivation

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Images

Calamus unspecified picture

Distribution

Calamus world distribution map, present in Angola, Andorra, Australia, Benin, Bangladesh, Central African Republic, China, Congo, Fiji, Gabon, Ghana, Guinea, Gambia, Guinea-Bissau, Indonesia, Lao People's Democratic Republic, Liberia, Moldova (Republic of), Myanmar, Nigeria, Nepal, Philippines, Sudan, Senegal, Solomon Islands, Sierra Leone, Thailand, Taiwan, Province of China, Uganda, Viet Nam, Vanuatu, Samoa, and Zambia

Identifiers

LSID urn:lsid:ipni.org:names:325996-2
WFO ID wfo-4000005953
COL ID 8VVV2
BDTFX ID 98596
INPN ID 446577
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Calamus Cornera Ceratolobus Zalaccella Rotang Rotanga Calospatha Daemonorops Schizospatha

Lower taxons

Calamus castaneus Calamus dasyacanthus Calamus maturbongsii Calamus pachypus Calamus pholidostachys Calamus suaveolens Calamus temii Calamus wanggaii Calamus wuliangshanensis Calamus bankae Calamus essigii Calamus acanthophyllus Calamus acanthospathus Calamus adspersus Calamus aidae Calamus altiscandens Calamus angustifolius Calamus anomalus Calamus arborescens Calamus arugda Calamus ashtonii Calamus asperrimus Calamus bacularis Calamus balingensis Calamus baratangensis Calamus barbatus Calamus batanensis Calamus bicolor Calamus billitonensis Calamus bimaniferus Calamus boniensis Calamus bousigonii Calamus brandisii Calamus burckianus Calamus burkillianus Calamus caesius Calamus calapparius Calamus caryotoides Calamus ceratophorus Calamus ciliaris Calamus cockburnii Calamus concinnus Calamus conjugatus Calamus convallium Calamus corneri Calamus crinitus Calamus deerratus Calamus delicatulus Calamus densiflorus Calamus depressiusculus Calamus dianbaiensis Calamus diepenhorstii Calamus dilaceratus Calamus dioicus Calamus discolor Calamus distentus Calamus divaricatus Calamus dongnaiensis Calamus elongatus Calamus erinaceus Calamus erioacanthus Calamus exilis Calamus filispadix Calamus fissus Calamus flabellatus Calamus flagellum Calamus floribundus Calamus formosanus Calamus foxworthyi Calamus gibbsianus Calamus godefroyi Calamus gonospermus Calamus grandifolius Calamus grandis Calamus griseus Calamus harmandii Calamus helferianus Calamus henryanus Calamus heteroideus Calamus hirsutus Calamus hookerianus Calamus inops Calamus insignis Calamus interruptus Calamus jenningsianus Calamus kandariensis Calamus karnatakensis Calamus lakshmanae Calamus laoensis Calamus latifolius Calamus lauterbachii Calamus laxissimus Calamus leiocaulis Calamus leptospadix Calamus leptostachys Calamus lewisianus Calamus lobbianus Calamus longipes Calamus longipinna Calamus longisetus Calamus longispathus Calamus macropterus Calamus macrosphaerion Calamus maiadum Calamus manan Calamus mattanensis Calamus megaphyllus Calamus melanacanthus Calamus melanochrous Calamus melanoloma Calamus melanorhynchus Calamus merrillii Calamus metzianus Calamus micracanthus Calamus micranthus Calamus microsphaerion Calamus minahassae Calamus minutus Calamus mirabilis Calamus mitis Calamus mogeae Calamus mollis Calamus moti Calamus muelleri Calamus multispicatus Calamus muricatus Calamus myriacanthus Calamus nagbettai Calamus nannostachys Calamus nanodendron Calamus nematospadix Calamus nicobaricus Calamus nielsenii Calamus occidentalis Calamus oligostachys Calamus optimus Calamus ornatus Calamus oxleyanus Calamus oxycarpus Calamus pachystemonus Calamus padangensis Calamus pandanosmus Calamus papuanus Calamus paspalanthus Calamus pedicellatus Calamus penicillatus Calamus peregrinus Calamus pilosissimus Calamus platyspathus Calamus plicatus Calamus poensis Calamus pogonacanthus Calamus poilanei Calamus polycladus Calamus polystachys Calamus pseudotenuis Calamus psilocladus Calamus pycnocarpus Calamus radiatus Calamus radicalis Calamus radulosus Calamus reinwardtii Calamus reticulatus Calamus reyesianus Calamus rhomboideus Calamus rhabdocladus Calamus rheedei Calamus ridleyanus Calamus robinsonianus Calamus rotang Calamus rudentum Calamus rumphii Calamus sabalensis Calamus samian Calamus sarawakensis Calamus schistoacanthus Calamus schlechterianus Calamus scipionum Calamus scleracanthus Calamus serrulatus Calamus simplex Calamus sordidus Calamus spectatissimus Calamus spinifolius Calamus spinulinervis Calamus stoloniferus Calamus tanakadatei Calamus tenompokensis Calamus tetradactyloides Calamus tetradactylus Calamus thwaitesii Calamus thysanolepis Calamus timorensis Calamus trachycoleus Calamus tumidus Calamus unifarius Calamus vattayila Calamus verticillaris Calamus vinosus Calamus viridissimus Calamus vitiensis Calamus walkeri Calamus whitmorei Calamus zollingeri Calamus zonatus Calamus beccarii Calamus evansii Calamus hukaungensis Calamus meghalayensis Calamus minor Calamus calospathus Calamus albidus Calamus sedens Calamus symphysipus Calamus sumbawensis Calamus acaulis Calamus bachmaensis Calamus centralis Calamus kontumensis Calamus lateralis Calamus nuichuaensis Calamus acanthochlamys Calamus cawa Calamus compsostachys Calamus comptus Calamus conirostris Calamus cumingianus Calamus cuthbertsonii Calamus didymocarpus Calamus didymophyllus Calamus draco Calamus egregius Calamus elegans Calamus equestris Calamus erectus Calamus filipendulus Calamus gracilipes Calamus gracilis Calamus halmaherensis Calamus klossii Calamus koordersianus Calamus korthalsii Calamus lambirensis Calamus leptopus Calamus macrochlamys Calamus malawaliensis Calamus manillensis Calamus marginatus Calamus modestus Calamus monticolus Calamus moorhousei Calamus moseleyanus Calamus moszkowskianus Calamus seriatus Calamus yentuensis Calamus parvulus Calamus orthostachyus Calamus perakensis Calamus pseudozebrinus Calamus rhytidomus Calamus ruber Calamus ruvidus Calamus sabensis Calamus salicifolius Calamus scabrispathus Calamus senalingensis Calamus toli-toliensis Calamus vestitus Calamus vidalianus Calamus spiralis Calamus ater Calamus badius Calamus banggiensis Calamus barfodii Calamus beguinii Calamus binnendijkii Calamus brachypodus Calamus brevicaulis Calamus bulubabi Calamus capillosus Calamus cheirophyllus Calamus cinereus Calamus clemensianus Calamus clivorum Calamus collarifer Calamus concolor Calamus dracunculus Calamus dumetosus Calamus erythrocarpus Calamus forbesii Calamus formicarius Calamus georgei Calamus heatubunii Calamus horridus Calamus ingens Calamus jacobsii Calamus johndransfieldii Calamus johnsii Calamus kampucheaensis Calamus katikii Calamus kebariensis Calamus komsaryi Calamus longispinosus Calamus longistipes Calamus lucysmithiae Calamus macrophyllus Calamus maculatus Calamus manii Calamus microstachys Calamus mollispinus Calamus nanduensis Calamus nicolai Calamus novae-georgii Calamus nuralievii Calamus ochrolepis Calamus ocreatus Calamus oligolepis Calamus oligophyllus Calamus oresbius Calamus oxycoccus Calamus papyraceus Calamus parutan Calamus parvifructus Calamus pintaudii Calamus plagiocyclus Calamus pogonotium Calamus politus Calamus posoanus Calamus pseudoconcolor Calamus quangngaiensis Calamus rarispinosus Calamus retroflexus Calamus ruptilis Calamus sabut Calamus sandsii Calamus sarasinorum Calamus sashae Calamus scapiger Calamus schlechteri Calamus sedisspirituum Calamus sekundurensis Calamus superciliatus Calamus tadulakoensis Calamus ursinus Calamus uschdraweitianus Calamus validus Calamus velutinus Calamus womersleyi Calamus wrightmyoensis Calamus zieckii Calamus melanacanthos Calamus baiyerensis Calamus confusus Calamus curranii Calamus flexilis Calamus hallierianus Calamus moorei Calamus collariferus Calamus scapigerus Calamus conspectus Calamus pumilus Calamus pedicellaris Calamus guruba Calamus wightii Calamus acamptostachys Calamus acehensis Calamus affinis Calamus asteracanthus Calamus cristatus Calamus croftii Calamus daemonoropoides Calamus eugenei Calamus kostermansii Calamus kunstleri Calamus lamprolepis Calamus loherianus Calamus longibracteatus Calamus luteus Calamus pachyrostris Calamus sepal Calamus serpentinus Calamus siberutensis Calamus singalanus Calamus spanostachys Calamus sparsiflorus Calamus spiculifer Calamus stenophyllus Calamus kurzianus Calamus manglaensis Calamus megalocarpus Calamus mogeanus Calamus oblatus Calamus pannosus Calamus nicolai Calamus schistocanthos Calamus viminalis Calamus balerensis Calamus takanensis Calamus treubianus Calamus unijugus Calamus urdanetanus Calamus fertilis Calamus geniculatus Calamus heteracanthus Calamus jenkinsianus Calamus karuensis Calamus kiahii Calamus kingianus Calamus kjellbergii Calamus macrorhynchus Calamus palembanicus Calamus calicarpus Calamus andamanicus Calamus aruensis Calamus australis Calamus javensis Calamus niger Calamus tenuis Calamus ovoideus Calamus riedelianus Calamus siphonospathus Calamus subangulatus Calamus trichrous Calamus zebrinus Calamus gamblei Calamus axillaris Calamus oblongus Calamus pilosellus Calamus inermis Calamus glaucescens Calamus melanochaetes Calamus fissilis Calamus periacanthus Calamus depauperatus Calamus crassifolius Calamus digitatus Calamus holttumii Calamus leloi Calamus myrianthus Calamus nitidus Calamus paulii Calamus pygmaeus Calamus rugosus Calamus scabridulus Calamus usitatus Calamus warburgii Calamus zeylanicus