Fruit a schizocarp enclosed within the persistent calyx, shorter than the calyx, oblong, hard, glabrous, formed by 2 equal 1-seeded mericarps, ribbed on the outer surface, sometimes winged in the upper 1/3, minutely verruculose-asperulous on the seed part inside, separating from one another[a6].
Corolla white or yellowish, funnel-shaped or hypocrateriform; tube slender straight or curved, somewhat widened at the apex, glabrous outside; limb ± oblique, 5-lobed or 5-partite with lobes subequal or unequal, glabrous outside, papillose-hairy at the base as well as in the throat.
Calyx narrowly tubular, 5-angled, with a green longitudinal rib on each angle and hyaline whitish between the ribs, slightly oblique, 5-dentate or 5-apiculate, the teeth ± unequal; fruiting calyx dilating and splitting longitudinally at the front, from base to apex.
Ovary cylindric, glabrous, 1-carpellate, carpel 2-locular, each locule with one basal anatropous ovule; style terminal, filiform, glabrous; stigma ± oblique, divided at the apex into an anterior stigmatiferous lobe and a posterior small tooth.
Stamens 4, didynamous, included, inserted in the upper part of the corolla tube; filaments short, shortly glandular hairy; anthers ovate, basifixed.
Erect or spreading perennial herbs or undershrubs, ± woody at the base, ± densely pubescent, the [a4] indumentum often with gland-tipped hairs.
Leaves opposite-decussate, simple, entire, serrate or dentate, lobed or pinnatipartite, sessile or petiolate.
Flowers sessile or shortly pedicellate in terminal spikes or racemes, slightly zygomorphic[a5].
Seeds linear, without albumen.