Pandanus Parkinson

Screwpine (en)

Genus

Angiosperms > Pandanales > Pandanaceae

Characteristics

Trees with candelabra-like branching and spiny adventitious roots; stem with almost annular scars of leaf-bases. Leaves with acute or attenuate apex, armed with small spines on margins, midrib beneath and sometimes on the twin lateral ventral pleats; midrib keeled beneath; secondary longitudinal veins numerous and clearly visible; tertiary transverse veins visible or obscure. Inflorescences terminal, capitate or spicate; bracts armed on the margins and sometimes on the midrib beneath, the lower peduncular ones green and attenuate at the apex, the more distal ones coloured and acute; flowers sessile, without a perianth. Male flowers in spicately arranged spadices; stamens many; filaments free or connate; anthers 2-thecous, oblong or linear, apiculate or obtuse. Female flowers in spicate or racemosely arranged bracteate spadices, or in a solitary spadix, without staminodes. Ovaries free or connate, each with a solitary ovule; placenta sub-basal but parietal. Fruit woody or drupaceous, ovoid, ellipsoid or spherical, often somewhat trigonous; carpels with a ± free stigma-remnant (pileus), this pyramidal, rounded or almost flat and only separated from other carpel-tips by sulci; style absent or vestigial; stigma (in species of Flora area) rounded-cordate to hippocrepiform or reniform, ± as long as broad, plane to oblique or sometimes vertically oriented, the basal groove adaxial (in 1-locular drupes) or centroscopic (in more-locular phalanges); pericarp thin, glabrous; upper mesocarp fibrous and with spongy pith, with several hollows; lower mesocarp fibrous-fleshy; endocarp thinly ligneous to thick and bony, whitish to dark brown, each locule furnished with a ‘tube’ of fibers running upward to the underside of the stigma, and a basal opening (through which germination occurs). Seed erect, the testa membranous; placenta lateral, slightly fibrous; endosperm firm.
More
Dioecious, terrestrial, semi-aquatic or rarely epiphytic (not in Australia) trees or shrubs, sometimes stoloniferous, often with aerial or prop roots with well-developed root caps. Leaves crowded at top of branches, arranged in 3 spirals, amplexicaul at base (in Australia), linear-ensiform, M-shaped in T.S. Male inflorescence branched (in Australia), with large bracts. Male flowers: stamens numerous, usually in umbel-like or racemose arrangements on stemonophores, sometimes inserted directly on rachis. Female inflorescences consisting of 1 or more globular, ellipsoid or cylindric heads (cephalia), nearly hidden within large bracts when flowering. Female flowers of 1–several carpels, each usually uni-ovulate; styles absent or spiniform; stigmas variously shaped. Fruiting heads globular, ellipsoidal or cylindric, of numerous crowded but free drupes or 1 or more polydrupes (phalanges). Endocarp woody. Seed coat thin, membranous. Seeds germinating within fruit. See also Stone (1993: 435), Green (1994: 415). [This description from the Flora of Australia Volume 39 treatment of Pandanaceae by Wilson (2011) may require revision to exclude any elements of the more recently recognised genus Benstonea that has been segregated from Pandanus (see Callmander et al. 2012)-Editor, 21 October 2021.]
Trees, shrubs, or herbs, evergreen, dioecious. Stems simple or branched, erect or prostrate, often with stiltlike, verrucose prop roots and aerial roots, sometimes virtually absent, often suckering. Leaves simple, terminal, sessile, densely arranged in corkscrew spirals, 3-or 4-seriate, linear, often spinose-serrate on margin and on keeled midvein abaxially. Inflorescence of large, bracteate heads or spadices; perianth absent. Male inflorescence paniculate with spiciform branches subtended by secondary, usually colored, spathes, branches covered with numerous stamens; flowers not individually distinguishable. Female inflorescence of globose to cylindric clusters or several-carpelled aggregates; flowers not individually distinguishable; carpels 1-ovuled; staminodes absent in female flower; placentation subbasal. Fruit a hard drupe, syncarpous, comprising an aggregation of individual connate, angled, fibrous phalanges; mesocarp sometimes hollow; exocarp fleshy; endocarp woody or bonelike; locules 1 or more; phalanges separating at maturity; stigma persistent, capitate. Seed solitary.
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Hardiness (USDA) 9-12

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Cultivation

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Images

Pandanus unspecified picture
Pandanus unspecified picture

Distribution

Pandanus world distribution map, present in Angola, Andorra, Australia, Benin, Bangladesh, Bermuda, Central African Republic, China, Congo, Cook Islands, Comoros, Cabo Verde, Costa Rica, Cuba, Fiji, Gabon, Ghana, Guinea, Gambia, Guinea-Bissau, Indonesia, Jamaica, Kenya, Lao People's Democratic Republic, Liberia, Moldova (Republic of), Madagascar, Maldives, Mali, Myanmar, Mozambique, Mauritius, Norfolk Island, Nigeria, Nepal, Nauru, Pitcairn, Philippines, Puerto Rico, Réunion, Sudan, Senegal, Solomon Islands, Sierra Leone, Somalia, Seychelles, Togo, Thailand, Tokelau, Tonga, Tuvalu, Taiwan, Province of China, Tanzania, United Republic of, Uganda, United States of America, Viet Nam, Vanuatu, Wallis and Futuna, Samoa, Yemen, and Zambia

Identifiers

LSID urn:lsid:ipni.org:names:31599-1
WFO ID wfo-4000027868
COL ID 8W23Q
BDTFX ID -
INPN ID 446269
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Barrotia Pandanus

Lower taxons

Pandanus maromokotrensis Pandanus polyglossus Pandanus scopula Pandanus scortechinii Pandanus saxatilis Pandanus semiarmatus Pandanus spinifer Pandanus spinistigmaticus Pandanus sparganioides Pandanus spathulatus Pandanus sibuyanensis Pandanus pyramidalis Pandanus quadrifidus Pandanus prainii Pandanus prostratus Pandanus pseudobathiei Pandanus radicans Pandanus princeps Pandanus rabaiensis Pandanus rollotii Pandanus rostratus Pandanus rotumaensis Pandanus retusus Pandanus verecundus Pandanus variabilis Pandanus vitiensis Pandanus yuleensis Pandanus vinaceus Pandanus coriaceus Pandanus zamboangensis Pandanus toliarensis Pandanus wiehei Pandanus sulcatus Pandanus subglobosus Pandanus teuszii Pandanus thomensis Pandanus tonkinensis Pandanus pinensis Pandanus yandeensis Pandanus lifouensis Pandanus pistikos Pandanus gracilialatus Pandanus mesos Pandanus ouveaensis Pandanus yoshioi Pandanus mareensis Pandanus latiloculatus Pandanus tenuimarginatus Pandanus pukapukaensis Pandanus stipiformis Pandanus subcylindricus Pandanus veillonii Pandanus verticalis Pandanus recavisaxosus Pandanus bryanii Pandanus daenikeri Pandanus induratus Pandanus adpressus Pandanus cuneiformis Pandanus efateensis Pandanus minimus Pandanus ngunaensis Pandanus planatus Pandanus radifer Pandanus spissus Pandanus anomesos Pandanus bilinearis Pandanus brongniartii Pandanus cavatus Pandanus nogarete Pandanus malgrasii Pandanus kajui Pandanus lacuum Pandanus lorencei Pandanus albifrons Pandanus analamerensis Pandanus silvanus Pandanus burmanicus Pandanus sulawesicus Pandanus decastigma Pandanus iwen Pandanus freetownensis Pandanus glaphyros Pandanus peekelii Pandanus mussauensis Pandanus isis Pandanus kaviengensis Pandanus longissimus Pandanus neomecklenburgensis Pandanus novibritannicus Pandanus rabaulensis Pandanus bismarckensis Pandanus parvicentralis Pandanus gabonensis Pandanus barbellatus Pandanus sylvicola Pandanus ambohitantelensis Pandanus flagellibracteatus Pandanus isalicus Pandanus majungensis Pandanus angolensis Pandanus bilobatus Pandanus gossweileri Pandanus raynalii Pandanus chevalieri Pandanus liberiensis Pandanus akeassii Pandanus lachaisei Pandanus amaryllifolius Pandanus alpestris Pandanus altissimus Pandanus aldabraensis Pandanus aggregatus Pandanus acaulescens Pandanus buinensis Pandanus brookei Pandanus boivinii Pandanus borneensis Pandanus brachus Pandanus brachyspathus Pandanus carrii Pandanus ceratostigma Pandanus calcis Pandanus calostigma Pandanus candelabrum Pandanus calceiformis Pandanus capitellatus Pandanus capusii Pandanus canaranus Pandanus aprilensis Pandanus aquaticus Pandanus aragoensis Pandanus aristatus Pandanus ambongensis Pandanus analamazaotrensis Pandanus ananas Pandanus apoensis Pandanus biliranensis Pandanus bicornis Pandanus barai Pandanus basalticola Pandanus balenii Pandanus linguiformis Pandanus leram Pandanus ligulatus Pandanus longicaudatus Pandanus livingstonianus Pandanus loherianus Pandanus marinus Pandanus manensis Pandanus marginatus Pandanus magnicavernosus Pandanus longipedunculatus Pandanus longipes Pandanus longissimepedunculatus Pandanus longistylus Pandanus maevaranensis Pandanus lamekotensis Pandanus involutus Pandanus julianettii Pandanus imerinensis Pandanus hystrix Pandanus iceryi Pandanus christmatensis Pandanus immersus Pandanus incertus Pandanus koordersii Pandanus krauelianus Pandanus labyrinthicus Pandanus lais Pandanus kaida Pandanus kanehirae Pandanus klossii Pandanus paludosus Pandanus palustris Pandanus panayensis Pandanus oligocephalus Pandanus platyphyllus Pandanus piniformis Pandanus polyacris Pandanus peyrierasii Pandanus penetrans Pandanus petrosus Pandanus pentodon Pandanus pendulinus Pandanus megacarpus Pandanus menicostigma Pandanus mindanaensis Pandanus microcarpus Pandanus microcephalus Pandanus nemoralis Pandanus neoleptopodus Pandanus nitidus Pandanus nobilis Pandanus darwinensis Pandanus gracilis Pandanus grallatus Pandanus graminifolius Pandanus gladiifolius Pandanus decipiens Pandanus dyeri Pandanus echinodermops Pandanus edulis Pandanus elatus Pandanus dorystigma Pandanus distentus Pandanus fibrosus Pandanus depauperatus Pandanus erectus Pandanus exaltatus Pandanus exiguus Pandanus eydouxia Pandanus dinagatensis Pandanus cookii Pandanus corniferus Pandanus clausus Pandanus connatus Pandanus clandestinus Pandanus comatus Pandanus heterocarpus Pandanus hooglandii Pandanus validus Pandanus umbonatus Pandanus heterostigma Pandanus mac-gregorii Pandanus castaneus Pandanus sandakanensis Pandanus pseudolais Pandanus busuangaensis Pandanus marojejicus Pandanus kuepferi Pandanus insuetus Pandanus sechellarum Pandanus schoddei Pandanus serratus Pandanus scopulorum Pandanus scabrifolius Pandanus salailuaensis Pandanus sambiranensis Pandanus samoensis Pandanus sarasinorum Pandanus sphaeroideus Pandanus spicatus Pandanus spondiophyllus Pandanus stellatus Pandanus stelliger Pandanus spiralis Pandanus simplex Pandanus singaporensis Pandanus sinicola Pandanus sphaerocephalus Pandanus solms-laubachii Pandanus solomonensis Pandanus pugnax Pandanus pulcher Pandanus punctulatus Pandanus pungens Pandanus pristis Pandanus rigidifolius Pandanus radula Pandanus ramosii Pandanus rapensis Pandanus reclinatus Pandanus rex Pandanus reticulatus Pandanus rubricinctus Pandanus welwitschii Pandanus vogelensis Pandanus viscidus Pandanus whitmeeanus Pandanus yasawaensis Pandanus yirrkalaensis Pandanus yvanii Pandanus kimlangii Pandanus manamboloensis Pandanus tenuifolius Pandanus tenuipedunculatus Pandanus taveuniensis Pandanus subacaulis Pandanus subulorum Pandanus sumatranus Pandanus unicornutus Pandanus turritus Pandanus tutuilaensis Pandanus unguifer Pandanus tsaratananensis Pandanus medialis Pandanus pistos Pandanus semipilaris Pandanus gibberosus Pandanus fidelis Pandanus pritchardiae Pandanus crassilix Pandanus laticonvexus Pandanus noumeaensis Pandanus amicalis Pandanus tongatapuensis Pandanus vavauensis Pandanus puniceus Pandanus arrectialatus Pandanus dasodes Pandanus recavilapideus Pandanus rectus Pandanus spheniskos Pandanus multidentatus Pandanus amnicola Pandanus crenifer Pandanus globatus Pandanus alticonvexus Pandanus breviendocarpicus Pandanus quinarius Pandanus onesuaensis Pandanus parou Pandanus pyramidos Pandanus metaceus Pandanus echinops Pandanus muralis Pandanus cissei Pandanus sikassoensis Pandanus farakoensis Pandanus andringitrensis Pandanus pluriaculeatus Pandanus tabellarius Pandanus alveolatus Pandanus serpentinicus Pandanus amissus Pandanus tenuiflagellatus Pandanus ambalavaoensis Pandanus leuconotus Pandanus dictyotus Pandanus associatus Pandanus nanofrutex Pandanus nervosus Pandanus regalis Pandanus satabiei Pandanus arapepe Pandanus sierraleonensis Pandanus oblongicapitellatus Pandanus triangularis Pandanus granulosus Pandanus alifer Pandanus gazelleensis Pandanus vandamii Pandanus denudatus Pandanus arenicola Pandanus tubulatus Pandanus tolanarensis Pandanus latistigmaticus Pandanus bemarahensis Pandanus membranaceus Pandanus cuneatus Pandanus pseudochevalieri Pandanus laferrerei Pandanus crassicollis Pandanus tiassaleensis Pandanus parachevalieri Pandanus guineabissauensis Pandanus djalonensis Pandanus insolitus Pandanus abbiwii Pandanus columellatus Pandanus brevifrugalis Pandanus pitcairnensis Pandanus calvus Pandanus senegalensis Pandanus alatus Pandanus alkemadei Pandanus aecuatus Pandanus acladus Pandanus acanthostylus Pandanus chiliocarpus Pandanus polycephalus Pandanus brevistipes Pandanus brosimos Pandanus brassii Pandanus bullii Pandanus burkillianus Pandanus butayei Pandanus calamianensis Pandanus botryoides Pandanus brachycarpus Pandanus brachyphyllus Pandanus bracteosus Pandanus calcicola Pandanus calathiphorus Pandanus cephalotus Pandanus ceylanicus Pandanus cheilostigma Pandanus camarinensis Pandanus caudatus Pandanus caricosus Pandanus carmichaelii Pandanus archboldianus Pandanus aridus Pandanus boemiensis Pandanus assamensis Pandanus atropurpureus Pandanus apiculatus Pandanus antaresensis Pandanus augustianus Pandanus beccarii Pandanus biceps Pandanus basilocularis Pandanus basedowii Pandanus bipollicaris Pandanus bipyramidatus Pandanus balfourii Pandanus austrosinensis Pandanus bakeri Pandanus balansae Pandanus bathiei Pandanus bantamensis Pandanus barkleyi Pandanus levuensis Pandanus leptocaulis Pandanus lamprocephalus Pandanus limbatus Pandanus lateralis Pandanus leptopodus Pandanus laxespicatus Pandanus leiophyllus Pandanus leptocarpus Pandanus malgassicus Pandanus mangokensis Pandanus mapola Pandanus longecuspidatus Pandanus martellii Pandanus maximus Pandanus macrophyllus Pandanus luzonensis Pandanus irregularis Pandanus boninensis Pandanus intricatus Pandanus kaernbachii Pandanus johorensis Pandanus joskei Pandanus lanutooensis Pandanus inundatus Pandanus huynhii Pandanus ijzermanni Pandanus juliferus Pandanus kedahensis Pandanus kajewskii Pandanus kamiae Pandanus kerchovei Pandanus kinabaluensis Pandanus kirkii Pandanus mc-keei Pandanus katatonos Pandanus pallidus Pandanus paloensis Pandanus pancheri Pandanus papuanus Pandanus paracalensis Pandanus mayotteensis Pandanus odorifer Pandanus oligocarpus Pandanus palawensis Pandanus orculiformis Pandanus ornatus Pandanus oviger Pandanus philippinensis Pandanus piricus Pandanus platycarpus Pandanus pleiocephalus Pandanus pluriloculatus Pandanus penicillus Pandanus patelliformis Pandanus patina Pandanus penangensis Pandanus perrieri Pandanus pervilleanus Pandanus moalaensis Pandanus monophalanx Pandanus multibracteatus Pandanus micracanthus Pandanus microstigma Pandanus militaris Pandanus multifurcatus Pandanus obeliscus Pandanus oblatus Pandanus obovatus Pandanus obsoletus Pandanus novohibernicus Pandanus occultus Pandanus navicularis Pandanus multispicatus Pandanus muricatus Pandanus myriocarpus Pandanus namakiensis Pandanus obconicus Pandanus multidrupaceus Pandanus fruticosus Pandanus dauphinensis Pandanus daymanensis Pandanus galeatus Pandanus galorei Pandanus gemmifer Pandanus gilbertanus Pandanus furcatus Pandanus flagellaris Pandanus dasystigma Pandanus floribundus Pandanus forbesii Pandanus forsteri Pandanus foveolatus Pandanus hata Pandanus guillaumetii Pandanus halleorum Pandanus gillespiei Pandanus hemisphaericus Pandanus dammannii Pandanus decumbens Pandanus glauciferus Pandanus glaucocephalus Pandanus grusonianus Pandanus emarginatus Pandanus diversus Pandanus dyckioides Pandanus ensifolius Pandanus drupaceus Pandanus dubius Pandanus durio Pandanus dolichopodus Pandanus diffusus Pandanus decus-montium Pandanus esculentus Pandanus fanningensis Pandanus conoideus Pandanus clementis Pandanus houlletii Pandanus hermaphroditus Pandanus helicopus Pandanus concretus Pandanus concinnus Pandanus conglomeratus Pandanus concavus Pandanus columnaris Pandanus conicus Pandanus cyaneoglaucescens Pandanus hermsianus Pandanus clarkei Pandanus crinifolium Pandanus hermesii Pandanus cubicus Pandanus cumingianus Pandanus nusbaumeri Pandanus kabaenaensis Pandanus daitoensis Pandanus ankaranensis Pandanus palakkadensis Pandanus callmanderiana Pandanus humbertii Pandanus caricosus Pandanus sermollianus Pandanus taluucensis Pandanus tectorius Pandanus tsingycola Pandanus bernardii Pandanus beguinii Pandanus benstoneoides Pandanus halmaherensis Pandanus sermolliana Pandanus lejophyllus Pandanus belepensis Pandanus korwae Pandanus mangalorensis Pandanus montanus Pandanus letocartiorum Pandanus rheophilus Pandanus pweleensis Pandanus tazoanii Pandanus fetosus Pandanus paucicarpellatus Pandanus mauricei Pandanus bowersiae Pandanus brevicornutus Pandanus mammillaris Pandanus kusaicola Pandanus discostigma Pandanus cupribasalis Pandanus pygmaeus Pandanus alveatus Pandanus horizontalis Pandanus lageniformis Pandanus oblongus Pandanus vandermeeschii Pandanus rechingerii Pandanus reineckei Pandanus reticulosus Pandanus pseudomontanus Pandanus macrostigma Pandanus navigatorum Pandanus cornerii Pandanus pseudocollinus Pandanus robinsonii Pandanus jaffrei Pandanus humilior Pandanus fusinus Pandanus faviger Pandanus purpurascens Pandanus tectorius Pandanus urophyllus Pandanus utilis Pandanus sylvestris