Englerina Tiegh.

Shrubs 0.5–2 m., with a single haustorial attachment, glabrous or shortly hairy; hairs simple or slightly branched, spreading, with an understorey of minute irregular trichomes, often giving a scurfy appearance. Leaves opposite or subopposite, the lowermost on flowering branchlets sometimes intergrading with bud-scales, sessile to generally petiolate; lamina sometimes coriaceous, but characteristically thin, sometimes flushed coppery brown, penninerved, but generally 1–several lower nerves more ascending. Flowers in pedunculate umbels, 2–20, often standing up like candles from horizontal branches; bract unilaterally developed from a generally shallow cupular base, not or scarcely exceeding the calyx, often slightly gibbous. Calyx annular or cup-shaped, sometimes lacerate or irregularly 4-lobed. Corolla 4-lobed, joined for one-fifth to two-thirds, red, yellow, orange or pink and white, often darker on bud-swellings.. Buds generally 4-angular, with vents marked by bosses, often with a slight to distinctly flask-shaped basal swelling, generally at least slightly swollen over the anthers, apiculate, obtuse, truncate or 4-bossed at the tip; lobes erect, reflexed or revolute, linear to distinctly spathulate above, sometimes hardened and shiny on the inner face of the dilated part, sometimes widened at the base around vents; tube split unilaterally, the V-slit extending halfway to almost the base, often papillate inside, at least along the sutures. Stamens attached at or near the base of corolla-lobes, the adnate part sometimes swollen or continued as a marked ridge down the corolla-tube, the free part tapered upwards, involute or inflexed at anthesis, produced into a tooth in front of the anther; anthers 4-thecous, with the connective slightly produced, minutely bifid. Style filiform, 4-angular; stigma obovoid to globular. Berry urceolate to obovoid, with persistent calyx, generally red; seed, where known, orange or yellow.