Gaultheria L.

Snowberry (en), Gaulthérie (fr)

Genus

Angiosperms > Ericales > Ericaceae

Characteristics

Evergreen shrubs or half-shrubs, mostly terrestrial, rarely epiphytic, erect or rarely procumbent, prostrate or creeping. Leaves spirally arranged or very rarely opposite, mostly serrate. Racemes simple or compound, i.e. forming panicles, whether or not leafy, perulate (i.e. with basal bracts) or practically eperulate, axillary or terminal, sometimes reduced to few-flowered fascicles or a solitary flower. Flowers rather small. Pedicel sustained by a large basal bract and 2(-3) higher inserted, mostly small bracteoles. Calyx 5-partite to various degree, lobes ± imbricate, persistent, becoming enlarged and swollen (and succulent) in fruit, finally rarely white or pinkish, mostly dark purple or almost blackish. Corolla campanulate or urceolate, shortly 5-lobed, lobes spreading or recurved, ± imbricate. Stamens 10 (in a few extra-Mal. spp. 8), included, attached around the disk to the base of the corolla; filaments mostly dilated at or above the base; anthers dorsifix, introrse, each cell produced at the apex into a very short and often broad tubule which opens by an apical ± oblique pore or slit, and whose back wall mostly is bidentate or biaristate apically. Disk ± 10-lobed, glabrous. Ovary superior or semi-inferior, 5-ribbed, 5-celled, with thick, bilobed axile, ∞-ovuled placentas; style simple, columnar; stigma small, ± obtuse. Capsule thin-to thick-walled, 5-celled, ± globose, dry, mostly rather completely included by the enlarged and ± succulent calyx, loculicidally dehiscent from the apex into 5 valves. Seeds ∞, minute, subglobose, angled; testa reticulate; albumen carnose; embryo cylindric.
More
Shrubs or subshrubs, (sometimes rhizomatous or stoloniferous and rooting at nodes). Stems erect or procumbent; twigs glabrous or hairy. Leaves persistent, aromatic; blade ovate, elliptic, or orbiculate to subcordate or reniform, coriaceous, margins serrate, crenate, or ciliate, plane or revolute, surfaces glabrous or hairy; venation reticulodromous or brochidodromous. Inflorescences axillary, racemes, 2-12-flowered, sometimes flowers solitary; (bracteoles closely subtending flowers). Flowers: sepals (4-)5, connate basally to nearly their entire lengths, (sometimes exceeding petals), ovate, deltate, or cordate; petals (4-)5, connate ca. 1/2 to nearly their entire lengths, white or cream to pink, corolla urceolate to campanulate, lobes much shorter than tube; stamens 8 or 10, included, (inserted at base of ovary); filaments straight, flattened, usually widest proximally, glabrous or hairy, sometimes papillose, without spurs; anthers with 2-4 awns or without awns, dehiscent by pores with ventral slits, (white disintegration tissue present dorsally along connective); pistil 4-5-carpellate; ovary 5-locular; stigma truncate or capitate. Fruits capsular, 5-valved, globose, fleshy, (surrounded by persistent, fleshy calyx). Seeds 20-80+, ovoid; testa smooth. x = 11, 12, 13.
Upright terrestrial or epiphytic shrubs, prostrate undershrubs or rarely small trees. Leaves persistent, short petiolate, usually serrate, pinnately veined. Inflorescences terminal panicles, axillary racemes or rarely solitary in the axils of the leaves; pedicels clearly articulated or jointed below the flower; bracteoles 2 or several, basal, medial or closely subtending the pedicels. Flowers with the calyx (4)5-parted, usually accrescent and becoming somewhat fleshy; corolla campanulate to urceolate, white to pinkish or reddish; stamens (8)10, the filaments distinct, basally expanded, borne on the base of the corolla, the anthers with white tissue on the abaxial side and often with 2, ascendent, terminal awns on each anther sac which open introrsely by a single apical pore; stigma obtuse or truncate, the style straight, the ovary superior or rarely partly inferior, (4)5-locular, ovules 5-10 (or many). Fruit a (4)5-valved, loculicidal capsule en-closed by the usually colored, slightly to conspicuously fleshy calyx and thus sometimes appearing superficially berrylike; seeds usually numerous, small, an-gled, and wingless.
Shrubs evergreen. Stems erect, creeping, or procumbent. Leaves spirally arranged, petiolate; leaf blade serrate or rarely entire. Flower usually 5-merous, sometimes 4-merous, in axillary or terminal racemes or panicles, or solitary; bracteoles variable in position. Calyx deeply divided. Corolla usually white, urceolate, campanulate, or tubular, shallowly lobed. Stamens included; filaments flattened, usually dilated towards base; anthers oblong, dehiscing by terminal pores, with 2–4 awns or minute projections. Ovary superior or semi-inferior, with many ovules per locule. Stigma truncate. Calyx at fruiting accrescent, fleshy; capsule dehiscing loculicidally or sometimes irregularly [fruit a berry]. Seeds small, unwinged.
Fls 4–5-merous; cal closely subtended by 2 bracteoles, campanulate to saucer-shaped, deeply divided; cor tubular to campanulate, shallowly lobed; stamens included; filaments short, flat; anthers oblong, the pollen-sacs separate or nearly so, each tipped by 2 erect awns; ovary 4–5-locular, wholly or partly superior; style short, columnar; stigma truncate; capsule thin-walled, completely enclosed in the fleshy, white or colored, accrescent cal, forming a dry or mealy berry-like fr with a characteristic flavor; erect to creeping shrubs with alternate, persistent lvs and usually white fls in racemes or panicles or (in our spp.) solitary in or just above the axils. 150, widespread.
Evergreen, erect or prostrate shrubs, occasionally tree-like. Lvs usually alternate, rarely opposite, usually toothed; petiole short or occasionally 0. Fls in terminal or axillary racemes or panicles, less commonly solitary, (4)-5-merous, pedicellate, with bracts and bracteoles. Calyx lobed, usually deeply so, accrescent. Corolla lobed, urceolate to campanulate. Stamens 10, included; anthers with 2 or 4 awns or obtuse. Disc 10-lobed. Ovary glabrous. Fr. a 5-valved capsule invested by enlarged and fleshy calyx and thus berry-like, rarely calyx persistent but dry (5 N.Z. spp.). Seeds numerous, minute.
Calyx 5-lobed, persistent; corolla 5-lobed, urceolate to campanulate, with 5 imbricate lobes. Stamens 10; anthers 2-celled, not exserted, dehiscing by pores, each cell tipped by 2 erect awns; filaments ± dilated at base. Ovary 5-loculed, style cylindric, stigma simple. Capsule loculicidally dehiscent, often invested by enlarged and succulent calyx; seeds ∞, minute. Widespread genus with c. 120 spp., mostly shrubs. The N.Z. spp. endemic, except G. depressa, which occurs in Tasmania.
Life form perennial
Growth form shrub
Growth support -
Foliage retention evergreen
Sexuality hermaphrodite
Pollination -
Spread -
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Flower color
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Fruit color
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Nitrogen fixer -
Photosynthetic pathway -

Environment

The genus is nearly exclusively terrestrial, much in contrast to the predominantly epiphytic genus Diplycosia. Most spp. occur as a substage in mountain forests, or in open country scrub, especially on mountain tops.
Light -
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Hardiness (USDA) 4-10

Usage

Uses. Many species contain in all their parts a volatile oil of peculiar odour, the so-called „winter-green-oil" (methyl-salicylate), which is greatly in repute against rheumatism. The smell becomes apparent or stronger after crushing the leaves or flowers. From the Malesian species G. punctata is used in the first place, known as „gondopuro" or „gandapura" to the Javanese. Less used are G. leucocarpa or G. nummularioides; from several other species in Malesia a strong smell has been stated by the col-lectors, but no use has been reported. The leaves are chewed occasionally together with „sirih" or used as a medicinal tea. The fruits are edible, but apparently not eaten by man.
Uses medicinal oil tea
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 30 - 60
Germination temperacture (C°) 18 - 21
Germination luminosity light
Germination treatment stratification
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Optimum temperature (C°) -
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Images

Gaultheria unspecified picture

Distribution

Gaultheria world distribution map, present in Australia, China, Hong Kong, Japan, Sri Lanka, New Zealand, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:30002565-2
WFO ID wfo-4000015402
COL ID 4MGL
BDTFX ID 86468
INPN ID 192723
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Gaultheria Brossaea

Lower taxons

Gaultheria akaensis Gaultheria bryoides Gaultheria abbreviata Gaultheria adenothrix Gaultheria amoena Gaultheria anastomosans Gaultheria antarctica Gaultheria antipoda Gaultheria atjehensis Gaultheria barbulata Gaultheria bracteata Gaultheria buxifolia Gaultheria caespitosa Gaultheria codonantha Gaultheria colensoi Gaultheria depressa Gaultheria dumicola Gaultheria x fagifolia Gaultheria foliolosa Gaultheria glaucifolia Gaultheria glomerata Gaultheria gracilescens Gaultheria griffithiana Gaultheria hapalotricha Gaultheria heteromera Gaultheria hirta Gaultheria hispida Gaultheria hispidula Gaultheria humifusa Gaultheria insana Gaultheria insipida Gaultheria kemiriensis Gaultheria lanceolata Gaultheria lanigera Gaultheria leucocarpa Gaultheria longibracteolata Gaultheria losirensis Gaultheria macrostigma Gaultheria malayana Gaultheria megalodonta Gaultheria minuta Gaultheria mundula Gaultheria myrtilloides Gaultheria oppositifolia Gaultheria oreogena Gaultheria ovatifolia Gaultheria paniculata Gaultheria parvula Gaultheria phillyreifolia Gaultheria poeppigii Gaultheria pullei Gaultheria purpurascens Gaultheria purpurea Gaultheria pyrolifolia Gaultheria pyroloides Gaultheria reticulata Gaultheria rupestris Gaultheria salicifolia Gaultheria santanderensis Gaultheria schultesii Gaultheria sclerophylla Gaultheria semi-infera Gaultheria serrata Gaultheria x serrulata Gaultheria seshagiriana Gaultheria sleumeri Gaultheria sleumeriana Gaultheria steyermarkii Gaultheria strigosa Gaultheria subcorymbosa Gaultheria taiwaniana Gaultheria tasmanica Gaultheria tenuifolia Gaultheria tetramera Gaultheria trichophylla Gaultheria vaccinioides Gaultheria vernalis Gaultheria wardii Gaultheria longiracemosa Gaultheria notabilis Gaultheria novaguineensis Gaultheria nubicola Gaultheria nubigena Gaultheria nummularioides Gaultheria pumila Gaultheria rigida Gaultheria setulosa Gaultheria solitaria Gaultheria tomentosa Gaultheria trigonoclada Gaultheria acuminata Gaultheria cardiosepala Gaultheria celebica Gaultheria chiriquensis Gaultheria dialypetala Gaultheria dolichopoda Gaultheria domingensis Gaultheria elegans Gaultheria fragrantissima Gaultheria howellii Gaultheria hypochlora Gaultheria x intermedia Gaultheria itatiaiae Gaultheria japonica Gaultheria x jordanensis Gaultheria eriophylla Gaultheria cuneata Gaultheria suborbicularis Gaultheria alnifolia Gaultheria marginata Gaultheria borneensis Gaultheria appressa Gaultheria arfakana Gaultheria gracilis Gaultheria crassa Gaultheria hookeri Gaultheria leschenaultii Gaultheria stereophylla Gaultheria straminea Gaultheria sinensis Gaultheria erecta Gaultheria racemulosa Gaultheria brevistipes Gaultheria acroleia Gaultheria angustifolia Gaultheria thymifolia Gaultheria lohitiensis Gaultheria berberidifolia Gaultheria bradeana Gaultheria jingdongensis Gaultheria pernettyoides Gaultheria praticola Gaultheria prostrata Gaultheria pseudonotabilis Gaultheria rengifoana Gaultheria ulei Gaultheria viridiflora Gaultheria myrsinoides Gaultheria mucronata Gaultheria procumbens Gaultheria shallon