Blechnum L.

Midsorus fern (en), Blechne (fr), Blechnum (fr)

Genus

Pteridophytes > Polypodiales > Blechnaceae > Blechnoideae

Characteristics

Terrestrial and lithophytic ferns. In Malesia none is persistently epiphytic. Some species develop an above-ground woody caudex up to 2 m high and may form extensive colonies interconnected in the early stages of development either by underground rhizomes or by above-ground stoloniferous connections. Scales on the rhizome are essentially similar to those at the base of the stipe; those on other parts of the plant in some species show a range of forms but are rarely clathrate; hairs are present on some species. The sterile lamina is typically pinnate, pinnatifid, or pinnatisect, rarely entire and if so usually in a juvenile stage of a plant’s development; veins in the majority are free, single, or dichotomising. Heteroblasty (for the succession of sterile fronds) is characteristic for some species. The fertile lamina in many species has the pinnae much reduced in width and the elongate linear sori may almost completely cover the abaxial surface (the ‘lomarioid’ condition). In other species the fertile pinnae appear the same as the sterile, on the adaxial surface. On the abaxial face the linear sori are closely aligned to the costa occupying a narrow region of the pinna undersurface (the ‘eublechnoid’ condition); intermediate arrangements are occasionally to be found in some taxa. An indusium is present in all taxa. Spores are monolete, bilateral and show a range of exine and perispore ornamentation.
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Small to medium terrestrial, lithophytic or, rarely, epiphytic ferns. Rhizome creeping, erect (sometimes forming a small trunk) or scandent. Fronds usually dimorphic but sterile and fertile fronds similar in some species. Lamina usually pinnate, pinnatisect or pinnatifid, but simple or lobed in some species; margins entire to serrate; veins free, usually forked and often terminating in glands close to the margin. Sori continuous, on either side of costa, ± covering the lower surface in dimorphic species with narrow fertile pinnae, or as a narrow longitudinal strip on either side of the costa in species with broad fertile pinnae; indusium present. Spores with variable surface ornamentation.
Plants terrestrial, moderate-sized or large. Rhizome usually erect or ascending, stout, dictyostelic, densely scaly; scales dark brown to brown, glossy, lanceolate, entire, basifixed. Fronds clustered, long stipitate; stipe robust, scaly at base, above naked; lamina pinnate, with reduced auricular lower pinnae, leathery; pinnae entire, linear, margin entire or serrulate; terminal pinna entire, similar to lateral pinnae; veins free, parallel, simple or forked. Sori linear, forming long coenosori adjacent and parallel to costa; indusium attached to commissure, facing toward costa; annulus longitudinal, interrupted, of 14-28 cells. Spores elliptic, usually smooth, with perispore.
Terrestrial, occasionally lithophytic ferns. Rhizome, stipe and sometimes pinna midvein with abaxial, lanceolate to linear-lanceolate medium to dark brown or black scales; scales sometimes attached to tubercles which remain to give a rough texture if scales are shed. Young fronds occasionally red. Fronds monomorphic or dimorphic; fertile fronds usually erect with narrower more widely spaced pinnae than the often prostrate sterile fronds; pinna margin sharply toothed; texture chartaceous to harsh; veins forked, at least in fertile pinnae. Sori borne on anastomoses parallel to pinna midvein.
Plants terrestrial or rarely on rock. Stems creeping to ascending or erect, slender to stout, sometimes climbing [rarely subarborescent]; scales brown or black. Leaves monomorphic or variously dimorphic, cespitose to scattered. Blades pinnatifid to 1-pinnate, rarely simple or 2-pinnate. Rachis and costae glabrous, scaly, or hairy abaxially. Veins free, often forked. Sori borne on vascular commissures parallel to costae, 1 per side, normally uninterrupted, linear, continuous along length of costa. Spores with perine smooth to variously winged or rugose. x = 28, 29, 31, 32, 34, 36.
Sori elongate, || costa, one on each side; indusium opening towards costa, firm, elongate. Sporangia rather large, crowded; annulus vertical, incomplete. Spores bilateral, reniform to subglobose, us. smooth. Rhizome dictyostelic, paleate, erect or ascending, sts forming a distinct caudex; or rhizome creeping, sts scandent. Fronds us. pinnate to pinnatifid, us. coriac. Over 200 spp., mainly of the southern hemisphere.
Medium-sized, terrestrial ferns. Rhizome creeping. Fronds dimorphic. Lamina pinnate or pinnatifid; margins serrate or crenate; bulbils sometimes present in axils of uppermost pinnae. Sterile pinnae with anastomosing veins. Sori longitudinal, continuous on either side of costa, ±covering the lower surface of narrow fertile pinnae; indusium present.
Sori continuous along each side of the costa of a leaflet, a sterile lamina present beyond the sorus or not
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Growth form tree
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Root system rhizome
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Environment

A widespread genus with ± 180 species showing considerable diversity; abundant in temperate, warm temperate, subalpine and tropical-montane and some oceanic island habitats.
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Hardiness (USDA) 8-12

Usage

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Cultivation

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Images

Blechnum unspecified picture

Distribution

Blechnum world distribution map, present in Australia, China, Cayman Islands, Guatemala, Honduras, Indonesia, Sri Lanka, Mexico, New Zealand, Papua New Guinea, El Salvador, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:328189-2
WFO ID wfo-4000004744
COL ID 62HK5
BDTFX ID 86001
INPN ID 189907
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Austroblechnum Cranfillia Neoblechnum Oceaniopteris Lomaridium Orthogramma Sadleria Homophyllum Lomariobotrys Stegania Struthiopteris Bowringia Cleistoblechnum Parablechnum Blechnidium Spicanta Spicantopsis Steenisioblechnum Lomariocycas Spicanta ghiesbreghtii Blechnopsis Blechnum Diploblechnum Brainea Lomaria stolonifera Lomaria ghiesbreghtii Struthiopteris stolonifera Blechnopteris Lonchitis-aspera Pteridoblechnum Distaxia Mesothema Doodia Lomaria Austroblechnum stoloniferum Icarus

Lower taxons

Blechnum marquesense Blechnum acanthopodum Blechnum keysseri Blechnum membranaceum Blechnum milnei Blechnum montanum Blechnum moorei Blechnum moritzianum Blechnum nesophilum Blechnum norfolkianum Blechnum norfolkianum Blechnum novae-zelandiae Blechnum puniceum Blechnum revolutum Blechnum x rodriguezii Blechnum rosenstockii Blechnum rufum Blechnum ryanii Blechnum vulcanicum Blechnum norfolkense Blechnum difforme Blechnum dilatatum Blechnum durum Blechnum filiforme Blechnum finlaysonianum Blechnum gemmascens Blechnum glaziovii Blechnum hieronymi Blechnum pallidum Blechnum palmiforme Blechnum pilosum Blechnum shaferi Blechnum sociale Blechnum tuerckheimii Blechnum wolamense Blechnum contiguum Blechnum moranianum Blechnum nigrum Blechnum castaneum Blechnum dissectum Blechnum doodianum Blechnum hindii Blechnum lineare Blechnum lyonii Blechnum marquesensis Blechnum maximum Blechnum medium Blechnum molle Blechnum neohollandicum Blechnum parrisiae Blechnum paschale Blechnum squarrosum Blechnum zeelandicum Blechnum longistipitatum Blechnum humbertii Blechnum divergens Blechnum fragile Blechnum insularum Blechnum ivohibense Blechnum inflexum Blechnum arcuatum Blechnum asperum Blechnum corralense Blechnum asplenioides Blechnum falciforme Blechnum lehmannii Blechnum schiedeanum Blechnum melanopus Blechnum lanceola Blechnum colombiense Blechnum diversifolium Blechnum christii Blechnum wardiae Blechnum minus Blechnum floresii Blechnum proliferum Blechnum dives Blechnum jamaicense Blechnum underwoodianum Blechnum auratum Blechnum tabulare Blechnum australe Blechnum punctulatum Blechnum xiphophyllum Blechnum binerve Blechnum lineatum Blechnum polypodioides Blechnum hastatum Blechnum microphyllum Blechnum mochaenum Blechnum penna-marina Blechnum schottii Blechnum gracile Blechnum confluens Blechnum cordatum Blechnum schomburgkii Blechnum chiriquanum Blechnum amabile Blechnum eburneum Blechnum opacum Blechnum sprucei Blechnum andinum Blechnum scaberulum Blechnum lima Blechnum fraseri Blechnum nudum Blechnum acutum Blechnum pteropus Blechnum loxense Blechnum auriculatum Blechnum dendrophilum Blechnum articulatum Blechnum socialis Blechnum rheophyticum Blechnum monomorphum Blechnum laevigatum Blechnum appendiculatum Blechnum austrobrasilianum Blechnum petiolare Blechnum capense Blechnum vestitum Blechnum venosum Blechnum reflexum Blechnum discolor Blechnum vieillardii Blechnum gracilipes Blechnum squamipes Blechnum colensoi Blechnum fernandezianum Blechnum stuebelii Blechnum organense Blechnum leyboldtianum Blechnum procerum Blechnum sampaioanum Blechnum usterianum Blechnum bicolor Blechnum bolivianum Blechnum bruneum Blechnum cochabambense Blechnum pazense Blechnum repens Blechnum smilodon Blechnum squamatum Blechnum vallegrandense Blechnum aequatoriense Blechnum ascendens Blechnum pinnatifidum Blechnum guayanense Blechnum squamosissimum Blechnum puberulum Blechnum nigrocostatum Blechnum obtusum Blechnum chambersii Blechnum fluviatile Blechnum gregsonii Blechnum wattsii Blechnum wurunuran Blechnum austrocaledonicum Blechnum spinulosum Blechnum capense Blechnum lherminieri Blechnum binervatum Blechnum raiateense Blechnum chauliodontum Blechnum corbassonii Blechnum hirsutum Blechnum decrescens Blechnum bonapartei Blechnum longipilosum Blechnum subcordatum Blechnum francii Blechnum brackenridgei Blechnum neglectum Blechnum reticulatum Blechnum melanocaulon Blechnum ludificans Blechnum leopoldense Blechnum fuscosquamosum Blechnum glabrescens Blechnum bakeri Blechnum polinesicum Blechnum patersonii Blechnum wagnerianum Blechnum cyatheoides Blechnum unisorum Blechnum longipinna Blechnum ambiguum Blechnum areolatum Blechnum nukuhivense Blechnum ensiforme Blechnum hancockii Blechnum obtusatum Blechnum werckleanum Blechnum rimbachii Blechnum flaccisquama Blechnum spannagelii Blechnum camfieldii Blechnum meridense Blechnum souleytianum Blechnum stipitellatum Blechnum caudatum Blechnum biforme Blechnum longicauda Blechnum oceanicum Blechnum integrifrons Blechnum longepetiolatum Blechnum madagascariense Blechnum cycadifolium Blechnum atropurpureum Blechnum werffii Blechnum cartilagineum Blechnum pacificum Blechnum gibbum Blechnum maxonii Blechnum lechleri Blechnum anthracinum Blechnum orientale Blechnum lenormandii Blechnum banksii Blechnum confusum Blechnum egregium Blechnum fullagari Blechnum geniculatum Blechnum howeanum Blechnum insigne Blechnum x pampasicum Blechnum triangularifolium Blechnum vittatum Blechnum malacothrix Blechnum brasiliense Blechnum attenuatum Blechnum magellanicum Blechnum occidentale Blechnum marginatum Blechnum spicant Blechnum spicant