Scleria P.J.Bergius

Nutrush (en)

Genus

Angiosperms > Poales > Cyperaceae

Characteristics

Monoecious (only the Australian S. sphacelata F.v.M. dioecious). Perennial. often stout herbs with short or creeping, often nodose rhizome, or annuals with fibrous, reddish roots. Stems solitary or more or less tufted, mostly erect, sometimes scrambling over bushes, trigonous or (in the Malesian spp.) triquetrous, leafy in the lower part or throughout, smooth or scabrid. Leaves 3-ranked, linear, sheathing the stem, smooth to very scabrous on the margins and the main nerves, the lower ones reduced to bladeless or almost bladeless sheaths; midrib prominent beneath, 2 lateral nerves prominent above; sheaths closed, not rarely 3-winged, eligulate, the apex on the ventral side truncate or produced into a tongue (contraligula). Inflorescence paniculate, consisting of a terminal partial panicle and usually some lateral ones, sometimes reduced to dense clusters, or glomerate-spiciform with glume-like bracts. Spikelets all bisexual, or bisexual and ♂, or ♀ and ♂; bisexual spikelets composed of 1 terminal ♀ flower and 1 to several lateral ♂ ones; ♀ spikelets with 1 ♀ flower and not rarely 1-2 lateral empty glumes (the reduced ♂ part); ♂ spikelets with several to numerous flowers. Glumes (except for the upper ones of the ♂ spikelets and of the ♂ part of the bisexual spikelets) distichous, in the lateral spikelets at right angles to the pertinent bract and prophyll, the lower 2-4 empty. Flowers unisexual, achlamydeous, the ♂ ones consisting of 1-3 stamens; anthers oblong to linear, with more or less produced connective; ♀ flowers with a 3-carpellate pistil; style continuous with the ovary, caducous, the base often persistent on the nut; stigmas 3. Nut globose, ovoid, ellipsoid, or pyramidal, terete or trigonous, smooth or variously sculptured, glabrous or hairy, with crustaceous pericarp, white, more rarely bluish, ultimately often discoloured, borne on a gynophore (cupula), which is dilated at the apex into a more or less trilobate, but sometimes much reduced, disk adhering to the ripe nut; outer cells of nut very small, quadrate-hexagonal.
More
Herbs usually perennial, sometimes annual, cespitose or not, rhizomatous or not. Culms solitary or not, unbranched proximal to inflorescence, trigonous, (6–)10–100(–125) cm (high-climbing into trees), weak or wiry. Leaves few to many per culm, cauline, 3-ranked; sheaths present; ligules usually well developed, sometimes obsolete; blades well developed (or rudimentary, sometimes absent on proximal leaves), shorter than, equaling, or overtopping culms, plane, keeled, or revolute, 0.5–9 mm wide, stiff, with prominent midvein, glabrous, scabrous, or pubescent. Inflorescences terminal, often axillary, sometimes pseudolateral, panicles, spikes, or 1 or more capitate or fasciculate clusters; spikelets 1–100+; bracts 1–3, ascending or erect, bristle-shaped, awl-shaped, or leaflike, sometimes appearing to be continuation of culm, 10–150 mm, shorter than or exceeding spikelets, glabrous or ciliate. Spikelets: scales 3–10+, each subtending flower, distichous, proximal 2–4 scales empty, proximal fertile scale pistillate, distal fertile scales staminate or sometimes empty; occasionally some spikelets entirely pistillate or staminate. Flowers unisexual; perianth absent; stamens 1–3; anthers 2–4 mm, apex usually mucronate or awned; styles linear, 2–3-fid, base deciduous or persistent, hypogynium rudimentary or prominent, slender or enlarged. Achenes trigonous or rounded, 1–4 mm, smooth, tuberculate, verrucose, or reticulate, occasionally pubescent.
Herbs, perennial or annual, rhizomatous or not; stolons sometimes present. Culms erect, 3-angled or rarely terete. Leaves cauline or both basal and cauline; sheaths at middle part of culm, winged or not, apex sometimes with contraligule opposite blade; leaf blade linear, scabrid, sheathing, usually prominently 3-veined. Involucral bracts leaflike and sheathing, setaceous, or glumelike; bractlets usually setaceous, rarely glumelike. Inflorescences terminal, paniculate, usually stout and elongated, sometimes reduced into a discontinuous spike. Unisexual spikelets usually dominant on inflorescence, rarely bisexual ones dominant; basal 2-4 glumes of spikelets empty. Male spikelets with several flowers. Female spikelets with a single flower. Bisexual spikelets with a basal female flower and several male flowers. Flowers unisexual. Male flowers: stamens 1-3; anthers linear or linear-oblong; connective apex often protruding and subulate. Female flowers: style base not inflated; stigmas 3. Nutlet brick red, white, or yellowish purple, spherical or ovoid, often obtusely 3-sided, smooth or variously sculptured, mostly shiny, glabrous or hairy (hairs simple or stellate), usually borne on a gynophore apically dilated into a ± 3-lobed disk.
Fls unisexual, the plant monoecious; spikelets small, the staminate few-fld, the pistillate with the lower scales empty and only the uppermost one fertile; perianth none; stamens 1–3; style trifid; achene bony or crustaceous, usually white or whitish, globose to ovoid or obscurely trigonous, obtuse or apiculate, in our spp. subtended by a simple or variously ornamented disk (hypogynium); sedges with trigonous culms and solitary or few, small, compact cymes. 200, warm reg.
Mostly leafy perennials, with small monoecious spikelets in terminal or axillary fascicles, or in interrupted spikes; fertile spikelets 1-flowered with several empty scales; staminate. spikelets many-flowered; achene crustaceous, globose to ovoid, frequently supported on a lobed or laciniate disc (hypogynium); stamens 1-3; style 3-cleft. About 100 species, chiefly in tropics of both hemispheres.
Life form perennial
Growth form herb
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Foliage retention deciduous
Sexuality
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Environment

As to altitude, the bulk of the species is found only in the tropical zone, below 1000 m, a few extend their area up to 1500 m, only one, S. terrestris, has been noted up to 2200 m; S. benthamii is only found at 1500 m.Most species seem to be indifferent to climate, but some are restricted to regions subject to an annual dry season, e.g. 4. S. brownii, 11. S. psilorrhiza, 12. S. junghuhniana, 28. S. tricuspidata, 29. S. laxa, and 32. S. pygmaeopsis.As to soil Sclerias are frequently found on sterile soils, and do not shun marshy lands, but on the whole they seem rather indifferent; S. cyathophora is, however, obviously silicicolous. Furthermore, S. poaeformis is a distinct swamp sedge, often forming tall, pure stands along lake shores.Though Sclerias occur in open places in the forest, they tend to be more common and gregarious in secondary growths and grasslands, along road-sides, on forest-edges, and in light forests. They have been frequently reported from Melaleuca woodland savannah. Almost invariably they take up a good portion of the stand, but we lack detailed figures; S. sumatrensis and S. biflora have been reported to be locally dominant.
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Hardiness (USDA) 8-12

Usage

Uses. The genus has hardly any economic value. The leaves of S. poaeformis are sometimes used for matting purposes, those of S. pergracilis as a medicine, and young plants of S. biflora are eaten with the rice as 'lalab'.
Uses medicinal
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Cultivation

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Images

Scleria unspecified picture

Distribution

Scleria world distribution map, present in Angola, Åland Islands, Andorra, Benin, Burkina Faso, Bangladesh, Bahamas, Belize, Bolivia (Plurinational State of), Botswana, Central African Republic, China, Congo, Comoros, Costa Rica, Cuba, Cayman Islands, Dominica, Ecuador, Eritrea, Ethiopia, Fiji, Micronesia (Federated States of), Gabon, Georgia, Ghana, Guinea, Gambia, Guinea-Bissau, Guatemala, Guyana, Honduras, Haiti, Indonesia, Jamaica, Japan, Kenya, Lao People's Democratic Republic, Liberia, Lesotho, Moldova (Republic of), Madagascar, Mali, Myanmar, Mozambique, Mauritania, Mauritius, Malaysia, Namibia, Nigeria, Nicaragua, Nepal, Panama, Peru, Philippines, Puerto Rico, Korea (Democratic People's Republic of), Paraguay, Réunion, Rwanda, Sudan, Senegal, Solomon Islands, Sierra Leone, Suriname, eSwatini, Seychelles, Chad, Togo, Thailand, Tonga, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Viet Nam, Wallis and Futuna, Samoa, Yemen, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:329988-2
WFO ID wfo-4000034707
COL ID 8W3GC
BDTFX ID -
INPN ID 627950
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Scleria Cylindropus Hymenolytrum Acriulus

Lower taxons

Scleria harlandii Scleria abortiva Scleria acanthocarpa Scleria adpressohirta Scleria alpina Scleria anceps Scleria andringitrensis Scleria angusta Scleria angustifolia Scleria anomala Scleria aromatica Scleria assamica Scleria atroglumis Scleria balansae Scleria bambariensis Scleria baroni-clarkei Scleria bequaertii Scleria boniana Scleria borii Scleria bourgeaui Scleria bracteata Scleria bradei Scleria brownii Scleria calcicola Scleria canescens Scleria carphiformis Scleria ciliaris Scleria ciliata Scleria clathrata Scleria colorata Scleria delicatula Scleria densispicata Scleria eggersiana Scleria elongatissima Scleria flexuosa Scleria georgiana Scleria glabra Scleria glomerulata Scleria hildebrandtii Scleria hirtella Scleria hispidior Scleria hispidula Scleria jiangchengensis Scleria kerrii Scleria khasiana Scleria lacustris Scleria latifolia Scleria laxa Scleria lithosperma Scleria lucentinigricans Scleria macrolomioides Scleria macrophylla Scleria madagascariensis Scleria martii Scleria melanotricha Scleria millespicula Scleria microcarpa Scleria minor Scleria mitis Scleria monticola Scleria muehlenbergii Scleria myricocarpa Scleria neocaledonica Scleria neogranatensis Scleria novae-hollandiae Scleria oligantha Scleria orchardii Scleria ovinux Scleria panicoides Scleria pauciflora Scleria paupercula Scleria pernambucana Scleria perpusilla Scleria pilosa Scleria pilosissima Scleria plusiophylla Scleria polycarpa Scleria polyrrhiza Scleria porphyrocarpa Scleria psilorrhiza Scleria rehmannii Scleria reticularis Scleria rugosa Scleria rutenbergiana Scleria scabriuscula Scleria scandens Scleria schenckiana Scleria schimperiana Scleria schliebenii Scleria scindens Scleria scrobiculata Scleria secans Scleria sellowiana Scleria sheilae Scleria skutchii Scleria sororia Scleria sphacelata Scleria spicata Scleria splitgerberiana Scleria sprucei Scleria stereorrhiza Scleria stipitata Scleria stipularis Scleria stocksiana Scleria suaveolens Scleria sumatrensis Scleria tenella Scleria tessellata Scleria testacea Scleria thwaitesiana Scleria transvaalensis Scleria tricuspidata Scleria triglomerata Scleria vaginata Scleria variegata Scleria venezuelensis Scleria verrucosa Scleria virgata Scleria vogelii Scleria warmingiana Scleria wrightiana Scleria zambesica Scleria achtenii Scleria amazonica Scleria annularis Scleria arcuata Scleria arguta Scleria baldwinii Scleria baronii Scleria benthamii Scleria bicolor Scleria buekiana Scleria bulbifera Scleria burchellii Scleria catophylla Scleria chevalieri Scleria chlorantha Scleria clarkei Scleria comosa Scleria composita Scleria cuyabensis Scleria cyathophora Scleria cyperina Scleria depauperata Scleria distans Scleria dregeana Scleria dulungensis Scleria erythrorrhiza Scleria fauriei Scleria filiculmis Scleria flagellum-nigrorum Scleria foliosa Scleria foveolata Scleria fulvipilosa Scleria gracillima Scleria greigiifolia Scleria guineensis Scleria havanensis Scleria hirta Scleria induta Scleria interrupta Scleria iostephana Scleria junghuhniana Scleria killipiana Scleria lagoensis Scleria leptostachya Scleria levis Scleria lingulata Scleria longispiculata Scleria macbrideana Scleria macrogyne Scleria melanomphala Scleria pachyrrhyncha Scleria natalensis Scleria naumanniana Scleria neesii Scleria nyasensis Scleria oblata Scleria obtusa Scleria oligochondra Scleria papuana Scleria parallella Scleria parvula Scleria patula Scleria pergracilis Scleria poeppigii Scleria poiformis Scleria procumbens Scleria purdiei Scleria purpurascens Scleria pusilla Scleria racemosa Scleria ramosa Scleria richardsiae Scleria robusta Scleria scabra Scleria schiedeana Scleria spiciformis Scleria swamyi Scleria tenacissima Scleria tepuiensis Scleria trialata Scleria tricholepis Scleria tryonii Scleria unguiculata Scleria verticillata Scleria veseyfitzgeraldii Scleria violacea Scleria williamsii Scleria woodii Scleria xerophila Scleria gaertneri Scleria afroreflexa Scleria brekiana Scleria biflora Scleria testacea Scleria goosenii Scleria rubrostriata Scleria chasmema Scleria pantadenia Scleria tricristata Scleria sobolifer Scleria didina Scleria aurantiaca Scleria bellii Scleria attenuatifolia Scleria chlorocalyx Scleria boivinii Scleria camaratensis Scleria corymbosa Scleria globonux Scleria goossensii Scleria hilsenbergii Scleria motleyi Scleria mutoensis Scleria pulchella Scleria radula Scleria remota Scleria robinsoniana Scleria schulzii Scleria uleana Scleria mucronata Scleria terrestris Scleria mikawana Scleria sieberi Scleria tonkinensis Scleria welwitschii Scleria huberi Scleria pooides Scleria castanea Scleria mackaviensis Scleria staheliana Scleria multilacunosa Scleria tropicalis Scleria laxiflora Scleria setulosociliata Scleria triquetra Scleria depressa