Lithocarpus Blume

Tanoak (en)

Genus

Angiosperms > Fagales > Fagaceae

Characteristics

Trees or shrubs; buttresses or stilt-roots occasionally present. Innovations densely yellowish brown to rufous tomentose by simple or stellate hairs. Terminal bud ovoid to ellipsoid; scales spirally imbricate. Stipules extrapetiolar, mostly caducous. Leaves spirally arranged, entire, thin-to thick-coriaceous, margin sometimes strongly recurved, glabrous or variously hairy at least beneath. Petiole terete or adaxially flat or furrowed. Inflorescence male, female, androgynous, or mixed, always erect, including bracts and bracteoles variously densely hairy. Male rachis solitary in the axil of a lower leaf or in a dense paniculate cluster on lateral or subterminal shoots, simple or much-branched; ♂ flowers solitary or in dichasial clusters of 3-7(-30) along the rachis, or in an androgynous inflorescence along the upper part of the rachis; perianth campanulate, (4-)6(-7)-lobed; stamens (8-)12(-15); filaments filiform, glabrous, anthers reniform, dorsifixed, 0.2-0.3(-0.5) mm long; pistillode always present, globose or subglobose, sometimes compressed, hairy. Female, androgynous or mixed rachis solitary in the axil of a higher leaf or on the upper part of the paniculate cluster; ♀ flowers solitary or in dichasial clusters of 3-7(-15) along the rachis, or in an androgynous inflorescence along the basal part of the rachis; perianth regularly 6-lobed, staminodes 10-12, rudimentary or well-developed and exceeding the perianth, sometimes polliniferous; style 3-6(-15), conical or terete, more or less connate or recurved, densely tomentose at base, stigma punctiform, terminal. Cupule solitary or in dichasial clusters of 3-7(-15) along the rachis, sessile or ½-2 cm stalked, enclosing a solitary fruit, cup-shaped, saucer-shaped to almost globular, variously lamellate, squamose, tuberculate or muricate, never truly spiny, never with vertical sutures; dehiscence none or irregular. Fruit in cross-section rounded, sometimes falsely multi-septate by intrusions of the endocarp; pericarp woody or bony, for the greater part free to adnate to the cupule, scar small to very large, glabrous or hairy. Cotyledons flat-convex. Germination hypogeal.
More
Trees or shrubs , evergreen. Terminal buds present, ovate, all scales imbricate. Leaves: stipules prominent on new growth, persistent around buds. Leaf blade leathery, margins entire or obscurely toothed to serrate, secondary veins unbranched, ± parallel, extending to margin. Inflorescences staminate and androgynous, axillary, often appearing terminal and branched by reduction of leaves, spicate, erect or ascending, rigid or flexible; androgynous inflorescences with pistillate cupules/flowers toward base and staminate flowers distally. Staminate flowers: sepals distinct; stamens 12(-18 or more) typically surrounding indurated pistillode covered with silky hairs. Pistillate flower 1 per cupule; sepals distinct; carpels and styles 3. Fruits: maturation in 2d year following pollination; cupule cup-shaped, without any indication of valves, covering proximal portion of nut, scaly, spines absent, scales strongly reflexed, hooked at tip; nut 1 per cupule, round in cross section, not winged. x = 12.
Trees or rarely shrubs, evergreen. Winter buds terminal, ovoid to ellipsoid, scales spirally imbricate. Stipules extrapetiolar. Leaves spirally arranged. Inflorescences male, female, or androgynous, in leaf axils toward base of branchlets or in a dense paniculate cluster on subterminal shoots, ± erect. Male inflorescences erect, simple or branched; flowers usually 3-5(-7) in dichasial clusters; perianth 4-6-lobed; stamens 10-12; rudimentary pistil small, enclosed by hairs. Female flowers solitary or in clusters of (2 or)3(-5), 1 or 2(or 3) well developed; perianth 6-lobed; staminodes 10-12; ovary 3(-6) loculed; styles (2 or)3(-5), (0.5-)1-2(-3) mm; stigmas a terminal pore. Cupules grouped together in cymes on rachis but often many aborted, corky, horny, woody, or crustaceous, completely or partly enclosing nut; bracts variously shaped. Nut 1 per cupule. Germination hypogeal; cotyledons flat-convex (although surface between cotyledons may not be completely flat).
Life form -
Growth form tree
Growth support -
Foliage retention evergreen
Sexuality monoecy
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

In Malesia the genus occurs mainly in the everwet lowland to montane forest, between 0-3000 m, more commonly below 1500 m, on various types of soil including limestone. In Central Java a few species have been recorded from pockets of everwet forest on Mts Dieng, Merapi, Ungaran, Muria, Lawu, and in E. Java on Mts Wilis, Kawi, Ardjuno, Tengger, and Idjen.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Lithocarpus unspecified picture

Distribution

Lithocarpus world distribution map, present in Argentina, Bangladesh, Bhutan, China, Spain, India, Japan, Korea (Republic of), Lao People's Democratic Republic, Moldova (Republic of), Myanmar, Nepal, Philippines, Thailand, Taiwan, Province of China, United States of America, and Viet Nam

Identifiers

LSID urn:lsid:ipni.org:names:30007732-2
WFO ID wfo-4000021973
COL ID 8VYYV
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Synaedrys Lithocarpus Pasania

Lower taxons

Lithocarpus ailaoensis Lithocarpus amherstianus Lithocarpus amoenus Lithocarpus amygdalifolius Lithocarpus apricus Lithocarpus areca Lithocarpus bennettii Lithocarpus bicoloratus Lithocarpus blumeanus Lithocarpus bonnetii Lithocarpus brachystachyus Lithocarpus brassii Lithocarpus brevicaudatus Lithocarpus bullatus Lithocarpus calolepis Lithocarpus calophyllus Lithocarpus cambodiensis Lithocarpus castellarnauianus Lithocarpus caudatifolius Lithocarpus caudatilimbus Lithocarpus celebicus Lithocarpus chittagongus Lithocarpus chiungchungensis Lithocarpus cinereus Lithocarpus cleistocarpus Lithocarpus coalitus Lithocarpus coinhensis Lithocarpus concentricus Lithocarpus confertus Lithocarpus confinis Lithocarpus confragosus Lithocarpus coopertus Lithocarpus corneri Lithocarpus corneus Lithocarpus cottonii Lithocarpus craibianus Lithocarpus crassifolius Lithocarpus cryptocarpus Lithocarpus cucullatus Lithocarpus cyrtocarpus Lithocarpus dalatensis Lithocarpus damiaoshanicus Lithocarpus daphnoideus Lithocarpus dinhensis Lithocarpus dodonaeifolius Lithocarpus echinifer Lithocarpus echinops Lithocarpus echinotholus Lithocarpus edulis Lithocarpus elaeagnifolius Lithocarpus elizabethiae Lithocarpus elmerrillii Lithocarpus encleisocarpus Lithocarpus falconeri Lithocarpus farinulentus Lithocarpus fenestratus Lithocarpus gymnocarpus Lithocarpus haipinii Lithocarpus indutus Lithocarpus irwinii Lithocarpus iteaphyllus Lithocarpus ithyphyllus Lithocarpus javensis Lithocarpus jenkinsii Lithocarpus jordanae Lithocarpus konishii Lithocarpus kontumensis Lithocarpus korthalsii Lithocarpus kostermansii Lithocarpus kozlovii Lithocarpus kunstleri Lithocarpus laetus Lithocarpus lampadarius Lithocarpus laoticus Lithocarpus laouanensis Lithocarpus leucodermis Lithocarpus levis Lithocarpus licentii Lithocarpus lindleyanus Lithocarpus listeri Lithocarpus litseifolius Lithocarpus longanoides Lithocarpus longinux Lithocarpus longipedicellatus Lithocarpus mianningensis Lithocarpus naiadarum Lithocarpus nantoensis Lithocarpus nieuwenhuisii Lithocarpus nodosus Lithocarpus oblanceolatus Lithocarpus oblancifolius Lithocarpus obovatilimbus Lithocarpus obscurus Lithocarpus ochrocarpus Lithocarpus oleifolius Lithocarpus pallidus Lithocarpus palungensis Lithocarpus philippinensis Lithocarpus pierrei Lithocarpus platycarpus Lithocarpus platyphyllus Lithocarpus pseudovestitus Lithocarpus pulcher Lithocarpus pusillus Lithocarpus pycnostachys Lithocarpus robinsonii Lithocarpus rosthornii Lithocarpus rotundatus Lithocarpus shunningensis Lithocarpus silvicolarum Lithocarpus skanianus Lithocarpus sogerensis Lithocarpus songkoensis Lithocarpus sphaerocarpus Lithocarpus tabularis Lithocarpus taitoensis Lithocarpus talangensis Lithocarpus tenuilimbus Lithocarpus tephrocarpus Lithocarpus uvariifolius Lithocarpus yongfuensis Lithocarpus vidalii Lithocarpus melanochromus Lithocarpus pseudoxizangensis Lithocarpus turbinatus Lithocarpus ovalis Lithocarpus ewyckii Lithocarpus fangii Lithocarpus glaber Lithocarpus xizangensis Lithocarpus variolosus Lithocarpus phansipanensis Lithocarpus proboscideus Lithocarpus stonei Lithocarpus arcaulus Lithocarpus beccarianus Lithocarpus clathratus Lithocarpus keningauensis Lithocarpus atjehensis Lithocarpus auriculatus Lithocarpus braianensis Lithocarpus chrysocomus Lithocarpus debaryanus Lithocarpus lithocarpaeus Lithocarpus echinulatus Lithocarpus eriobotryoides Lithocarpus aspericupulus Lithocarpus attenuatus Lithocarpus bacgiangensis Lithocarpus balansae Lithocarpus bassacensis Lithocarpus chevalieri Lithocarpus chienchuanensis Lithocarpus chifui Lithocarpus dasystachyus Lithocarpus fenzelianus Lithocarpus floccosus Lithocarpus fohaiensis Lithocarpus fordianus Lithocarpus formosanus Lithocarpus gaoligongensis Lithocarpus gigantophyllus Lithocarpus glutinosus Lithocarpus guinieri Lithocarpus hancei Lithocarpus handelianus Lithocarpus havilandii Lithocarpus henryi Lithocarpus himalaicus Lithocarpus honbaensis Lithocarpus howii Lithocarpus hypoglaucus Lithocarpus hystrix Lithocarpus imperialis Lithocarpus kawakamii Lithocarpus kemmaratensis Lithocarpus kingianus Lithocarpus kochummenii Lithocarpus lappaceus Lithocarpus lauterbachii Lithocarpus lepidocarpus Lithocarpus leptogyne Lithocarpus luteus Lithocarpus luzoniensis Lithocarpus lycoperdon Lithocarpus macilentus Lithocarpus magneinii Lithocarpus mairei Lithocarpus mariae Lithocarpus megacarpus Lithocarpus megalophyllus Lithocarpus megastachyus Lithocarpus meijeri Lithocarpus mekongensis Lithocarpus ollus Lithocarpus ombrophilus Lithocarpus oogyne Lithocarpus pachycarpus Lithocarpus pachylepis Lithocarpus rufescens Lithocarpus pachyphyllus Lithocarpus paihengii Lithocarpus pakhaensis Lithocarpus paniculatus Lithocarpus papillifer Lithocarpus pattaniensis Lithocarpus paviei Lithocarpus petelotii Lithocarpus porcatus Lithocarpus propinquus Lithocarpus psammophilus Lithocarpus pseudomagneinii Lithocarpus pseudomoluccus Lithocarpus pseudoreinwardtii Lithocarpus pseudosundaicus Lithocarpus qinzhouicus Lithocarpus quangnamensis Lithocarpus quercifolius Lithocarpus recurvatus Lithocarpus reinwardtii Lithocarpus revolutus Lithocarpus rhabdostachyus Lithocarpus rigidus Lithocarpus rouletii Lithocarpus rufovillosus Lithocarpus ruminatus Lithocarpus scyphiger Lithocarpus shinsuiensis Lithocarpus stenopus Lithocarpus suffruticosus Lithocarpus sulitii Lithocarpus sundaicus Lithocarpus syncarpus Lithocarpus thomsonii Lithocarpus toumorangensis Lithocarpus touranensis Lithocarpus triqueter Lithocarpus uraianus Lithocarpus urceolaris Lithocarpus vestitus Lithocarpus vidalianus Lithocarpus vinhensis Lithocarpus vinkii Lithocarpus wallichianus Lithocarpus woodii Lithocarpus xylocarpus Lithocarpus yersinii Lithocarpus echinophorus Lithocarpus elegans Lithocarpus erythrocarpus Lithocarpus gracilis Lithocarpus eucalyptifolius Lithocarpus harmandii Lithocarpus lucidus Lithocarpus eichleri Lithocarpus garrettianus Lithocarpus macphailii Lithocarpus maingayi Lithocarpus sootepensis Lithocarpus truncatus Lithocarpus trachycarpus Lithocarpus tubulosus Lithocarpus obovalifolius Lithocarpus perakensis Lithocarpus rassa Lithocarpus pseudokunstleri Lithocarpus rufus Lithocarpus scortechinii Lithocarpus solerianus Lithocarpus wrayi Lithocarpus polystachyus Lithocarpus sandakanensis Lithocarpus siamensis Lithocarpus harlandii Lithocarpus tenuinervis Lithocarpus hatusimae Lithocarpus hendersonianus Lithocarpus jacksonianus Lithocarpus jacobsii Lithocarpus tawaiensis Lithocarpus elephantum Lithocarpus ferrugineus Lithocarpus gougerotae Lithocarpus kalkmanii Lithocarpus kamengii Lithocarpus hallieri Lithocarpus magnificus Lithocarpus melataiensis Lithocarpus microbalanus Lithocarpus microlepis Lithocarpus mindanaensis Lithocarpus muluensis Lithocarpus nebularum Lithocarpus nhatrangensis Lithocarpus menadoensis Lithocarpus milroyi Lithocarpus moluccus Lithocarpus neorobinsonii Lithocarpus bentramensis Lithocarpus brochidodromus Lithocarpus burkillii Lithocarpus leiostachyus Lithocarpus bancanus Lithocarpus blaoensis Lithocarpus leiocarpus Lithocarpus leiophyllus Lithocarpus lemeeanus Lithocarpus aggregatus Lithocarpus andersonii Lithocarpus annamensis Lithocarpus annamitorus Lithocarpus acuminatus Lithocarpus apoensis Lithocarpus bolovenensis Lithocarpus campylolepis Lithocarpus cantleyanus Lithocarpus crassinervius Lithocarpus curtisii Lithocarpus carolinae Lithocarpus conocarpus Lithocarpus ducampii Lithocarpus echinocarpus Lithocarpus dolichostachys Lithocarpus longzhouicus Lithocarpus loratifolius Lithocarpus yangchunensis Lithocarpus glaucus Lithocarpus orbicarpus Lithocarpus eriobotryifolius Lithocarpus dahuoaiensis Lithocarpus vuquangensis Lithocarpus pulongtauensis Lithocarpus monticolus Lithocarpus submonticolus Lithocarpus parvulus Lithocarpus sericobalanos Lithocarpus cerifer Lithocarpus clementianus Lithocarpus cyclophorus Lithocarpus dealbatus Lithocarpus orbicularis