Stellaria L.

Starwort (en), Stellaire (fr), Stellaire au sens large (fr)

Genus

Angiosperms > Caryophyllales > Caryophyllaceae

Characteristics

Plants annual, winter annual, or perennial. Taproots usually slender, perennial taxa often rhizomatous, rooting at nodes. Stems prostrate to ascending or erect, simple or branched, terete or 4-angled. Leaves sometimes connate basally into sheath, often sessile; blade 1-veined, linear or lanceolate to ovate or deltate, succulent (S. crassifolia [gemmae], S. fontinalis, S. humifusa, and S. irrigua) or not, apex acute or obtuse. Inflorescences terminal, open cymes, rarely axillary (S. alsine, S. americana) or umbellate (S. umbellata), or terminal or axillary solitary flowers; bracts paired (1 in S. dicranoides), foliaceous, scarious and reduced, or absent. Pedicels erect, sometimes reflexed in fruit, glabrous or pubescent, not glandular. Flowers usually bisexual (S. dicranoides unisexual); perianth and androecium hypogynous or weakly perigynous; hypanthium cup-or disc-shaped; sepals (4-)5, distinct, green, occasionally purple tinged (S. irrigua) or red proximally (S. pallida), lanceolate to ovate-triangular, 2-12 mm, herbaceous (rarely coriaceous), margins often white, scarious, apex acute, acuminate, or obtuse, not hooded; petals (1-)5 or absent, white (sometimes translucent in S. borealis), not clawed, blade apex 2-fid usually for 3-5 its length (S. holostea occasionally laciniate); nectaries at base of filaments opposite sepals usually present, disc sometimes prominent; stamens (1-)5 or 10 or absent, arising from nectariferous disc (prominent in S. dicranoides and S. irrigua) at ovary base; filaments distinct; staminodes absent; styles [2-]3(-5), capitate to clavate, 0.2-7 mm, glabrous proximally; stigmas [2-]3(-5), terminal or subterminal, papillate (30×). Capsules globose to conic, opening by 3 or 6, occasionally 4, 8, or 10 ascending to recurved valves; carpophore present or absent. Seeds (1-)3-20+, yellow-brown to dark brown, globose to ellipsoid, laterally compressed, rarely shiny, papillate or rugose, rarely smooth, marginal wing absent, appendage absent. x = 10, 11, 12, 13, 15.
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Annual or perennial herbs, often much branched and clambering, the stems somewhat resilient and occasionally suffruticose below. Leaves opposite, petiolate or sessile and somewhat perfoliate, exstipulate, in Central America rather broad and membranaceous. Flowers in few-flowered dichasial cymes or less frequently solitary in the axils of slightly reduced leaves. Sepals 4-5, not connate. Petals 4-5, rarely absent, white, deeply 2-cleft. Stamens 4-10; anthers versatile, 2-celled, the flattened hypogynous filaments connate below forming a brief glandular disk. Ovary superior, carpels 2-4, with as many filiform styles free to the base; ovules 4-many, camnpylotropous on basal, central or free central placentae. Capsule deeply or shallowly dehiscing into as many 2-cleft or rarely entire valves as there are styles; seeds 4-many, cochleate, smooth or tuberculate, estrophiolate, the embryo curved about the perisperm.
Herbs annual, biennial, or perennial. Roots or rhizomes filiform and slender, roots sometimes robust, fleshy. Stems erect, ascending, or prostrate. Leaves compressed, variously shaped, rarely needlelike; stipules absent. Flowers terminal, seldom only axillary, in cymes or solitary, small; bracts immediately subtending calyx absent. Sepals (4 or)5. Petals (4 or)5, white, rarely green, usually 2-cleft nearly to base, rarely retuse or multilobed, sometimes absent. Stamens 2--5 or (6--)10. Ovary 1-loculed, rarely 3-loculed when young; ovules numerous, rarely several and only 1 or 2 mature; styles (2 or)3(or 4). Capsule orbicular or ovoid, opening by valves (1 or)2 × number of styles. Seeds (1 to) numerous, reniform, slightly compressed, tuberculate or smooth; embryo curved.
Small, annual or perennial herbs. Stems slender, often thickened at nodes, glabrous or hairy. Leaves exstipulate, sessile or petioled, glabrous or pubescent with simple or stellate hairs. Inflorescences in lax, dichotomous cymes. Flowers hypogynous or slightly perigynous. Sepals (4 or) 5, free, herbaceous, usually 3-nerved. Petals (4 or) 5, rarely ab-sent, white, usually bifid, almost to the base. Stamens (3-)5 or 10, rarely more or absent. Nectarial glands present in the form of a disc or separate glands between or inside the stamens. Ovary unilocular; styles 2 or 3. Fruit a capsule opening by 4-6 valves. Seeds mostly numerous, rarely few to 1, reniform or spherical, rugose or verrucose.
Herbs, annual or perennial, glabrous or sparsely hairy. Leaves opposite; stipules absent; floral leaves herbaceous to scarious. Flowers in spreading dichasia or partial monochasia, or solitary or 2-flowered, 5-merous, bisexual. Sepals 5, free, usually narrowly to broadly ovate. Petals usually 5, free, bifid, shorter to longer than sepals, in some taxa reduced or absent. Stamens 1–10. Staminodes 0–5. Ovary 1-celled; styles usually 3, free to base; placentation free-central. Fruit a conical capsule, splitting to middle or below, usually 6-valved (twice the number of styles). Seeds 2–many, subdiscoid to nautiloid to obloid; testa minutely to prominently tuberculate.
Annual or perennial herbs. Hairs usually sparse and eglandular, or 0. Lvs opposite, exstipulate, sessile or petiolate, glabrous or ciliate, linear, lanceolate, ovate or orbicular. Infl. a dichasium or reduced to 1-2 fls; bracts scarious to herbaceous; epicalyx 0. Sepals 5, free, not veined or (1)-3-veined. Petals white, divided almost to base, rarely 0. Stamens 10 or fewer. Styles 3. Fr. a capsule, usually ovoid, sometimes globose or cylindric, dehiscing by 6 teeth; carpophore 0. Seeds usually 4-10, ovoid, flattened, tuberculate or papillate, not winged.
Annual or perennial herbs, often with slender diffuse stems, glabrous or hairy. Leaves opposite, simple, entire, flat to (rarely) subulate, exstipulate. Inflorescences dichasial cymes which are terminal to the branches, rarely the flowers solitary. Sepals free, 4–5. Petals 4–5, more or less deeply bilobed, white, or occasionally absent. Stamens 5 + 5, or fewer by reduction. Styles generally 3 but sometimes 2, free. Capsule unilocular with 3 (or rarely 2) or 6 (rarely 4) valves. Seeds either numerous or 1–4.
Fls solitary in the forks of the stem, or in terminal cymes; sep 5; pet 5, bifid, often deeply so, or lacking; stamens mostly 10, sometimes fewer; styles mostly 3(4) but 5 in S. aquatica; ovules numerous; capsule ordinarily dehiscent by twice as many valves as there are styles, but in S. aquatica with 5 apically notched or shortly bifid valves; low annual or perennial herbs with mostly rather small, opposite, exstipulate lvs; × most commonly=13. (Alsine, Myosoton) 100+, widespread, esp. N. Temp.
Fls hermaphrodite, regular, solitary or in cymes. Sepals 4-5, free; petals white, 4-5, us. very deeply bifid, sts lacking; stamens 10 or fewer; ovary 1-celled, styles 3 (rarely 2 or 4-5), free to the base; capsule opening by twice as many teeth as there are styles. Slender annual to perennial herbs. Lvs opp., simple, entire, exstipulate. About 85 spp., cosmopolitan. The 6 spp. indigenous to N.Z. are all endemic. Several naturalized spp. are widespread weeds.
Leaves opposite, sessile or petiolate, simple, entire, flat or very rarely subulate, exstipulate.
Capsule usually more or less rounded, opening at the apex by 3 or 6 (rarely 2 or 4) valves.
Inflorescences of terminal dichasial cymes, rarely flowers solitary.
Annual or perennial herbs usually with slender diffuse stems.
Petals 4–5, white, deeply bilobed, or occasionally absent.
Seeds numerous or occasionally 1–4-, roundish-reniform.
Stamens 10 (5 + 5) or fewer by reduction.
Ovary 1-locular; styles (2) 3, free.
Sepals 4–5, free.
Life form
Growth form herb
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Sexuality hermaphrodite
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Root system rhizome tap-root
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Environment

Grows in a wide range of habitats in Australia including coastal dunes, heathlands, grasslands, herblands, swamps, fern communities in gully forests, underlying open eucalypt or Callitris woodlands, rainforests, alpine bogs, mallees and Chenopodiaceae or Leptospermum scrublands. The endemic species are often associated with wetter areas such as watercourses, areas that seasonally flood, drainage areas or dams. The weedy species are often associated with disturbed areas such as roadside drains, dams or cultivated areas. They can grow in a wide variety of soil types from sands to clays and on almost any rock type.
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Hardiness (USDA) 8-12

Usage

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Cultivation

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Images

Stellaria unspecified picture
Stellaria unspecified picture

Distribution

Stellaria world distribution map, present in Australia, China, New Zealand, and Panama

Identifiers

LSID urn:lsid:ipni.org:names:30001456-2
WFO ID wfo-4000036401
COL ID 4PZ
BDTFX ID 87263
INPN ID 197932
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Stellaria Mesostemma Cherleria Fimbripetalum

Lower taxons

Stellaria cuspidata Stellaria dichotoma Stellaria holostea Stellaria longipes Stellaria irazuensis Stellaria ovata Stellaria obtusa Stellaria venezuelana Stellaria uliginosa Stellaria umbellata Stellaria crassipes Stellaria elatinoides Stellaria bungeana Stellaria ciliatisepala Stellaria crassifolia Stellaria fennica Stellaria humifusa Stellaria hebecalyx Stellaria mannii Stellaria media Stellaria graminea Stellaria palustris Stellaria pallida Stellaria gyangtseensis Stellaria antoniana Stellaria aphanantha Stellaria vestita Stellaria chinensis Stellaria petiolaris Stellaria lanipes Stellaria gyirongensis Stellaria zangnanensis Stellaria tibetica Stellaria lanata Stellaria patens Stellaria souliei Stellaria parviumbellata Stellaria subumbellata Stellaria decumbens Stellaria arenarioides Stellaria pilosoides Stellaria concinna Stellaria montioides Stellaria webbiana Stellaria calycantha Stellaria monosperma Stellaria nipponica Stellaria fischerana Stellaria aphananthoidea Stellaria lanceolata Stellaria emirnensis Stellaria americana Stellaria apetala Stellaria corei Stellaria crispa Stellaria dicranoides Stellaria laevis Stellaria nitens Stellaria ruscifolia Stellaria parviflora Stellaria borealis Stellaria littoralis Stellaria radians Stellaria alsinoides Stellaria amblyosepala Stellaria brachypetala Stellaria martjanovii Stellaria persica Stellaria petraea Stellaria soongorica Stellaria winkleri Stellaria porsildii Stellaria poeppigiana Stellaria parva Stellaria reticulivena Stellaria nepalensis Stellaria wushanensis Stellaria henryi Stellaria pusilla Stellaria mainlingensis Stellaria depressa Stellaria yunnanensis Stellaria uda Stellaria congestiflora Stellaria ovatifolia Stellaria delavayi Stellaria aquatica Stellaria omeiensis Stellaria nubigena Stellaria alaskana Stellaria antillana Stellaria fontinalis Stellaria arvalis Stellaria chilensis Stellaria debilis d'urv. Stellaria cherleriae Stellaria discolor Stellaria ebracteata Stellaria filicaulis Stellaria hippoctona Stellaria imbricata Stellaria irrigua Stellaria turkestanica Stellaria longifolia Stellaria arisanensis Stellaria recurvata Stellaria dianthifolia Stellaria infracta Stellaria cryptopetala Stellaria pedersenii Stellaria weddellii Stellaria salicifolia Stellaria alaschanica Stellaria oxycoccoides Stellaria bistyla Stellaria strongylosepala Stellaria celsa Stellaria circinata Stellaria sennii Stellaria neglecta Stellaria nemorum Stellaria pubera