Xyris L.

Yelloweyed grass (en)

Genus

Angiosperms > Poales > Xyridaceae

Characteristics

Mostly perennial, paludose, grass-like herbs with fibrous roots; stembase very rarely thickened, often profusely producing shoots. Leaves basal, distichous on each shoot, ensiform, linear or filiform, sometimes twisted; sheaths with a membranous margin (in Mal.spp.) producing mucilage (?always), with or without a short ligule; limb glabrous or with numerous, small hard papillae, sometimes with a stout nerve in either margin. Flowers hermaphrodite, in terminal, few-to many-flowered heads, 3-merous, yellow to white, ephemeral, each in the axil of a conspicuous bract; bracts conchate, imbricate, spirally arranged, lower ones sterile; one to few flowers simultaneously in anthesis. Peduncles scape-like, terete to compressed, sometimes winged or ribbed, glabrous or with numerous hard papillae, at the base with some sheaths provided with a short limb. Bracts entire, ciliate, fimbriate or lacerate, with one complete main nerve and some complete or incomplete longitudinal secondary (descending) nerves, in the apical part mostly with a small minutely-papillose field. Calyx zygomorphic; lateral sepals navicular, with entire, dentate or ciliate crest, wings membranous, entire, glabrous or ciliate; median sepal membranous, spathel-liform or cap-shaped, enveloping the corolla, mostly obovate, 1-3(-5)-nerved, pushed out by the corolla in anthesis (?always). Corolla actinomorphic, ephemeral; petals with an orbicular to obovate limb and a long, narrow claw, free, cohering mutually or by the staminodes. Stamens mostly 3 fertile epipetalous inserted on the petals and 3 alternating staminodes, staminodes rarely absent, or all stamens fertile; filaments short; anthers basifix, dehiscing lengthwise extrorsely. Ovary superior, sessile to stipitate (in Australian spp. sometimes with 3 hard swellings at the top), 1-or 3-celled, or incompletely 3-celled. Placentas parietal, central, or basal, with ∞ ovules; styles filiform, apex 3-fid, stigmas mostly capitate. Fruit shape similar to that of the ovary but larger, loculicidally 3-valved. Seeds ellipsoid to obovoid, often ribbed, with a long funicle.
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Annual to perennial, scapose, rosulate, stems mostly short, cespitose, rarely rhizomatous. Leaves equitant, distichous, sheaths open, clasping, usually broad-based, with margins entire to ciliate or fimbriate, apically either narrowing gradually or abruptly to blade or there producing a ligule or auricle, blades mostly laterally compressed and 2-margined, mostly linear, less often thickened and ventrally sulcate to terete. Scapes single or few per shoot, arising from axillary buds, sheathed at base by a single, bladeless or bladed bract, terminating in a single spike (except in no. 3); spikes cone-like, chaffy-bracted, with 1 to many flowers; bracts in spiral or decussate, the lower 2-several empty, rarely forming a "leafy" involucre, commonly grading larger into the few to many flowering bracts, all or some with thicker central zone (dorsal area) of distinctly different texture and color or lacking dorsal area entirely. Sepals 3, generally hidden by subtending bract with 2 lateral, boat-shaped, free or connate, with variously margined keel and flanking a membranaceous inner (adaxial) sepal which enfolds the corolla in bud, detaching as the flower opens; petals 3, free (rarely connate), long-clawed, only the broad blades and claw tips exserted, these mostly yellow (in Africa some are blue); staminodes 3 (rarely lacking), mostly 2-branched with branches bearded; stamens mostly with flattened filaments and narrow, basifixed, introrse or laterally dehiscent anthers; ovary 1-or-3-locular, placentation marginal, parietal, basal, central or transitional, style tubular, slender, conduplicate, elongate, 3-branched just above petal claws, each branch terminating in a "U"-shaped stigma; stylar appendages absent. Capsules usually thinvalved; seeds (l-) few to many, mostly small and ovoid to fusiform, mostly longitudinally ribbed, opaque or translucent, embryo tiny, basal or lateral-basal.
Herbs, perennial, occasionally annual, rosulate. Stems simple, erect, sometimes caudiciform, short to branching, elongate. Leaves alternate, 2-ranked, equitant; blade mostly linear to filiform, flattened to nearly terete, margins smooth to variously papillate or scabrous. Inflorescences: scapes variously elongate; spike bracts imbricate, the proximal sterile, decussate, distal ones fertile, in spiral [decussate], bearing medially, usually subapically, a distinct and mostly differently textured and colored "dorsal area" or this indistinct or absent. Flowers solitary in bract axils; sepals 3, distinct, unequal, lateral (outer) sepals distinct [connate], chaffy, boat-shaped, chaffy, inner one membranous, capping corolla; keel papery, or scarious, variously winged, variously margined; petals 3, distinct [to connate], equal, strongly clawed; blade ephemeral, spreading, yellow to white; staminodes 3, distinct, strongly clawed, apically yoked, bearded with moniliform hairs (rarely reduced, beardless); stamens 3, epipetalous; anthers 2--4-locular; ovaries y thin-walled, placentation marginal to parietal [basal, free-central, axile]; styles elongate, tubular, 3-branched; stigmas 3, U-shaped or funnelform. Capsules 3-valved, usually thin, loculicidal. Seeds [1--]15--90 or more, mealy or translucent, ovoid to cylindric, 0.3--1(--4) mm, variously lined and cross-lined; endosperm starchy and proteinaceous; embryo basal-lateral, minute. x = 9.
Herbs perennial or rarely annual. Leaves equitant; leaf sheath sometimes with a short ligule; leaf blade linear to ensiform, smooth, papillose, rugose, or scaberulose. Peduncle scapelike, terete to compressed, sometimes winged or ribbed. Spikes few to many flowered; bracts brownish yellow to blackish brown, dull or shining, conchiform, rigid, abaxially minutely papillose toward apex when young, midvein conspicuous, margin scarious, entire, ciliate, fimbriate, or lacerate, apex emarginate, rounded, or acute. Sepals 3; lateral sepals with keel entire, dentate, or ciliate, wings membranous; median (anterior) sepal enclosing corolla or sometimes ± reduced or obsolete, thin and membranous, 1--3(--5)-veined. Petals 3, free; limb obovate; claw long and narrow. Stamens usually 3; filaments flattened; anthers basifixed, extrorse, dehiscing by vertical slits. Staminodes 3 or rarely absent, alternate with petals, often clawed, 2-cleft or 2-brachiate and branches with penicillate hairs. Ovary usually sessile, 1-or 3-loculed or imperfectly 3-loculed. Style filiform, apex 3-branched; stigmas usually U-shaped. Seeds obovoid to ellipsoid, longitudinally striate, apex usually apiculate.
Perennial, occasionally annual, tufted herbs of damp or wet situations. Leaves mostly basal, linear, terete, or lanceolate, sheathing at the base. Flowering scapes erect, simple, terminated by a solitary dense spike or head, and usually bearing 1 to several bladeless sheaths at the base. Flowering spikes or heads globose to elongate; bracts glumaceous, rather rigid or thin and papery, spirally imbricated, the lower usually sterile, the upper subtending the rather inconspicuous flowers. Flowers perfect, solitary and sessile in the axils of the bracts, yellow or blue, rarely white. Sepals 2-3, unequal. Petals 3, usually equal, obovate-spatulate, the basal claws free or united. Stamens 3, opposite the petals and adnate to claws; filaments usually short and flattened; anthers basifixed, 2-celled, dehiscent longi-tudinally. Staminodes (when present) 3, alternating with the petals, usually bifid at the tip, the branches usually tipped with brush-like tufts of moniliform hairs. Ovary 1-celled or imperfectly 3-celled; ovules numerous; style filiform, usually 3-cleft at the tip. Capsule loculicidal.
Annual or perennial herbs, often forming clumps; rhizomes, when present, short, horizontal or ascending. Leaves virtually all basal; ligule present or not. Inflorescence a condensed bracteate spike at the end of elongated peduncles which are often variously grooved or ridged; lower bracts of the spike sterile, occasionally elongated or reflexed. Flowers borne singly in the axils of the fertile bracts, short-lived; calyx of 3 unequal segments, the adaxial hooded and enclosing the flower-bud, the laterals asymmetric and often keeled, free from one another (sometimes fused basally in American species); corolla yellow (blue in at least one West African species). Stamens alternating with 3 usually bifurcate staminodes divided apically into many moniliform hairs. Ovary with parietal placentation (in East Africa). Fruit a loculicidal capsule.
Fls trimerous; anterior sep well developed, deciduous; pet distinct, with slender claw (hidden by the bract) and broad, yellow (white), toothed blade; staminodes present, bifid; placentas in most spp. (incl. ours) parietal, ± intruded distally; style-branches 3, alternate with the staminodes; seeds longitudinally lined. 250+, mainly warm reg.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
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Root system rhizome
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Nitrogen fixer -
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Environment

Restricted to marshy habitats, often on sandy, acid soils together with Cyperaceae, Eriocaulon, Dr oser a, Juncus, etc., and on borders of shallow swamps; locally they are often very common or even subgregarious. Single species are common in wet rice-fields, e.g. X. indica, which seems to be almost restricted to this anthropogenic habitat.Most spp. occur in the lowland, but several are bound to mountain marshes, e.g. X. ftabellata, X. grandis, and X. oreophila.
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Hardiness (USDA) 8-12

Usage

Uses -
Edible -
Therapeutic use -
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Cultivation

Mode -
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Images

Xyris unspecified picture

Distribution

Xyris world distribution map, present in Angola, Åland Islands, Andorra, Australia, Benin, Burkina Faso, Bangladesh, Belize, Bolivia (Plurinational State of), Brazil, Botswana, Central African Republic, China, Congo, Costa Rica, Cuba, Dominica, Ecuador, Ethiopia, Micronesia (Federated States of), Gabon, Georgia, Ghana, Guinea, Guinea-Bissau, Guatemala, Guyana, Honduras, Indonesia, Jamaica, Kenya, Lao People's Democratic Republic, Liberia, Lesotho, Moldova (Republic of), Madagascar, Mali, Myanmar, Mozambique, Mauritania, Mauritius, Malaysia, Namibia, Nigeria, Nicaragua, Nepal, Panama, Peru, Philippines, Palau, Puerto Rico, Paraguay, Rwanda, Sudan, Senegal, Sierra Leone, Suriname, eSwatini, Chad, Togo, Thailand, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Viet Nam, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:327087-2
WFO ID wfo-4000041064
COL ID 8W4S7
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Xyroides Xyris Ramotha

Lower taxons

Xyris byssacea Xyris almae Xyris anceps Xyris apureana Xyris aracamunae Xyris araracuarae Xyris archeri Xyris aristata Xyris bampsii Xyris blanchetiana Xyris brachyfolia Xyris brevifolia Xyris calderonii Xyris caparaoensis Xyris caroliniana Xyris columbiana Xyris complanata Xyris concinna Xyris consanguinea Xyris contracta Xyris correlliorum Xyris coutensis Xyris cyperoides Xyris diamantinae Xyris difformis Xyris dilatatiscapa Xyris dissitifolia Xyris disticha Xyris downsiana Xyris ekmanii Xyris elegantula Xyris eleocharoides Xyris elliottii Xyris erubescens Xyris esmeraldae Xyris exigua Xyris fibrosa Xyris flexifolia Xyris floridana Xyris formosana Xyris fuliginea Xyris gossweileri Xyris gracillima Xyris graminosa Xyris grandiceps Xyris graniticola Xyris grisebachii Xyris guaranitica Xyris guillenii Xyris hatschhachii Xyris hilariana Xyris huillensis Xyris humpatensis Xyris hystrix Xyris indivisa Xyris involucrata Xyris isoetifolia Xyris itambensis Xyris itatiayensis Xyris juncea Xyris juncifolia Xyris kradungensis Xyris kukenaniana Xyris lacerata Xyris laxiflora Xyris lejolyana Xyris leonensis Xyris lithophila Xyris lomatophylla Xyris longibracteata Xyris longiscapa Xyris macbrideana Xyris madagascariensis Xyris mallocephala Xyris mantuensis Xyris maxima Xyris melanopoda Xyris melanovaginata Xyris membranibracteata Xyris mentiens Xyris minarum Xyris metallica Xyris mexicana Xyris morii Xyris neblinae Xyris neocaledonica Xyris nigrescens Xyris nigricans Xyris nivea Xyris nubigena Xyris obcordata Xyris obscura Xyris obtusiuscula Xyris oligantha Xyris paleacea Xyris pallidula Xyris paradisiaca Xyris parvula Xyris pauciflora Xyris peteri Xyris phaeocephala Xyris picea Xyris pilosa Xyris piraquarae Xyris piresiana Xyris platylepis Xyris platystachya Xyris porcata Xyris regnellii Xyris reitzii Xyris retrorsifimbriata Xyris riparia Xyris roycei Xyris rubrolimbata Xyris rubromarginata Xyris savanensis Xyris schneeana Xyris serotina Xyris seubertii Xyris sparsifolia Xyris spathacea Xyris spectabilis Xyris spruceana Xyris stenocephala Xyris subasperula Xyris subglabrata Xyris subsetigera Xyris subtilis Xyris subulata Xyris tennesseensis Xyris toronoana Xyris torta Xyris tristis Xyris ustulata Xyris valdeapiculata Xyris valida Xyris veruina Xyris vestita Xyris villosicarinata Xyris affinis Xyris albescens Xyris coronata Xyris dawsonii Xyris harleyi Xyris ambigua Xyris amorimii Xyris andina Xyris angularis Xyris aquatica Xyris asperula Xyris atrovirida Xyris augusto-coburgii Xyris aurea Xyris bahiana Xyris baldwiniana Xyris bancana Xyris barteri Xyris bialata Xyris bicephala Xyris bissei Xyris blepharophylla Xyris boliviana Xyris borneensis Xyris brachysepala Xyris bracteata Xyris cachimbensis Xyris calcicola Xyris capensis Xyris capnoides Xyris carinata Xyris celiae Xyris chapmanii Xyris chimantae Xyris ciliata Xyris confusa Xyris congensis Xyris connosepala Xyris cryptantha Xyris cuatrecasana Xyris curassavica Xyris cylindrostachya Xyris dardanoi Xyris decussata Xyris densa Xyris diaphanobracteata Xyris dissimilis Xyris drummondii Xyris egleri Xyris exilis Xyris fallax Xyris ferreirae Xyris filifolia Xyris filiformis Xyris fimbriata Xyris flabelliformis Xyris foliolata Xyris frequens Xyris friesii Xyris frondosa Xyris fusca Xyris gerrardii Xyris glandacea Xyris globosa Xyris glochidiata Xyris goyazensis Xyris gracilis Xyris guianensis Xyris huberi Xyris hymenachne Xyris indica Xyris insignis Xyris intersita Xyris jataiana Xyris jolyi Xyris juncea Xyris jupicai Xyris kibaraensis Xyris kornasiana Xyris laevigata Xyris lagoinhae Xyris lanata Xyris laniceps Xyris lanuginosa Xyris lanulobractea Xyris laxifolia Xyris longifolia Xyris longisepala Xyris lucida Xyris luetzelburgii Xyris lugubris Xyris lutescens Xyris machrisiana Xyris makuensis Xyris marginata Xyris mello-barretoi Xyris mertensiana Xyris mexiae Xyris mima Xyris montana Xyris moraesii Xyris mucujensis Xyris natalensis Xyris nilssonii Xyris oblata Xyris ornithoptera Xyris paraensis Xyris pectinata Xyris plantaginea Xyris popeanus Xyris pranceana Xyris pratensis Xyris prolificans Xyris ptariana Xyris pterygoblephara Xyris pumila Xyris ramboi Xyris rigidiformis Xyris roraimae Xyris rubella Xyris scabrifolia Xyris sceptrifera Xyris schizachne Xyris schliebenii Xyris setigera Xyris sincorana Xyris smalliana Xyris sororia Xyris stenophylla Xyris stenophylloides Xyris stenostachya Xyris stricta Xyris subuniflora Xyris surinamensis Xyris symoensii Xyris tatei Xyris teinosperma Xyris tenella Xyris teres Xyris terrestris Xyris thysanolepis Xyris tomentosa Xyris tortilis Xyris tortula Xyris trachysperma Xyris tuberosa Xyris uleana Xyris uninervis Xyris unistriata Xyris vacillans Xyris vivipara Xyris wallichii Xyris wawrae Xyris witsenioides Xyris xiphophylla Xyris sulcatifolia Xyris nanuzae Xyris piranii Xyris kralii Xyris anamariae Xyris fredericoi Xyris bracteicaulis Xyris crassifunda Xyris submetallica Xyris guillauminii Xyris graomogolensis Xyris calostachys Xyris rehmannii Xyris velutina Xyris riopretensis Xyris rostrata Xyris scoparia Xyris pirapamae Xyris pulchella Xyris nervata Xyris nigra Xyris atrospicata Xyris cervii Xyris maparecida Xyris shepherdiana Xyris bituberosa Xyris buengkanensis Xyris thailandica Xyris aberdarica Xyris acrophila Xyris atrata Xyris capillaris Xyris imitatrix Xyris inaequalis Xyris liesneri Xyris linifolia Xyris lobbii Xyris navicularis Xyris porphyrea Xyris sanguinea Xyris straminea Xyris trachyphylla Xyris wurdackii Xyris grandis Xyris cipoensis Xyris consolida Xyris delicatula Xyris malmeana Xyris scabridula Xyris spinulosa Xyris teretifolia Xyris trichophylla Xyris welwitschii Xyris panacea Xyris aristida Xyris muelleri Xyris rupicola Xyris asterotricha Xyris atriceps Xyris labatii Xyris marojejyensis Xyris fugaciflora Xyris angustifolia Xyris anisophylla Xyris arachnoidea Xyris erosa Xyris bicarinata Xyris bicostata Xyris calostachys Xyris cheumatophila Xyris culmenicola Xyris ednae Xyris gongylospica Xyris kundelungensis Xyris kwangolana Xyris lacera Xyris neglecta Xyris operculata Xyris organensis Xyris oxylepis Xyris pancheri Xyris rhodolepis Xyris rigida Xyris sphaerocephala Xyris tasmanica Xyris decipiens Xyris festucifolia Xyris spathifolia