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Heads disciform or discoid, the outer fls in our spp. pistillate and fertile, the central ones perfect or sometimes sterile; invol bracts dry, imbricate, at least the inner scarious or with scarious margins; receptacle flat to convex or hemispheric, naked or beset with long hairs; style-branches flattened, truncate, penicillate; achenes ellipsoid or obovoid to nearly prismatic, scarcely compressed, usually glabrous; pappus none; herbs or shrubs, usually aromatic, with alternate, entire to dissected lvs and few to numerous small, ovoid to campanulate or hemispheric heads in a spiciform, racemiform, or paniculiform infl. 100+, N. Hemisphere and S. Amer.
Annual, biennial or perennial herbs or shrubs, taprooted or rhizomatous. Lvs usually pinnately divided, alternate. Capitula in terminal, pedunculate panicles, racemes or corymbs; involucral bracts in few rows, with membranous margins. Receptacle flat to hemispheric; scales 0, but pilose hairs sometimes present. Outer florets ♀ or ☿, tubular, brown, red, or yellow; inner florets ☿ or ♂, tubular. Achenes all similar, obovoid, subterete or flattened, smooth, faintly striate, or 2-ribbed; pappus 0 or a small scarious ring.
Annual or perennial herbs, subshrubs or shrubs, erect. Leaves entire, lobed or variously pinnatisect. Capitula commonly numerous per stem, disciform, pedunculate or sessile; involucre 2-or 3-seriate, with bracts gradational in length or not; receptacle epaleate. Outer florets female, fertile, 2–4-lobed. Disc florets bisexual (in Australia), sometimes functionally male; corolla 5-lobed. Achenes ±homomorphic, quadrangular, ±smooth or 2-ribbed, glabrous. Pappus absent.
