Herbs, annual or perennial, subshrubs, [trees], taprooted. Stems erect, usually much-branched, hairy, hairs stellate or 1-rayed and apparently simple. Leaves petiolate; stipules subpersistent, linear-lanceolate to lanceolate; blade ovate or broadly ovate to oblong-elliptic, lanceolate-elliptic, or ovate-elliptic, rhombic-ovate, or suborbiculate, often palmately 3–5-lobed, herbaceous, margins serrate to dentate, surfaces usually stellate-pubescent, palmately 3–7-veined. Inflorescences axillary or terminal, cymose [solitary flowers or fasciculate], 2 or 3(–5)-flowered, simple or ultimately 1–4-branched, clusters leaf-opposed at axils, sometimes appearing subspicate. Flowers: sepals 5, [triangular-lanceolate] narrowly oblong to linear, valvate, often cucullate, commonly with hornlike subapical apiculum, stellate-pubescent to glabrate; petals 5 [absent or relatively small], yellow, narrowly oblanceolate to oblong or linear-oblong, usually with basal, adaxial glands; stamens 5–25[–60], on androgynophore; staminodes 0; ovary 2–4[or 5]-locular; ovules 1 or 2 per locule; styles 1, simple, short-cylindric; stigmas 2–5-lobed. Capsules globose to ovoid-globose or ovoid, 2–3[–6]-valved, spiny, spines uncinate [straight], glabrous or hairy, indehiscent [dehiscence loculicidal]. Seeds 1–2 per locule, irregularly ovoid to obovoid or pyriform, smooth. x = 8.
Herbs, rarely subshrubs, annual or perennial, erect or procumbent, hairs with swollen base, stellate or with 1 ray and apparently simple. Leaves alternate, simple or palmately 3-5-lobed, palmately veined, margin serrate. Flowers solitary or arranged in small cymes or fascicles, axillary, opposite to leaves, bisexual; peduncle very short. Sepals 5, free, valvate, usually with hooked appendages at tip. Petals as many as sepals, free, thickened glands on base of adaxial surface. Stamens 5 to numerous; filaments free; anthers dorsifixed, subglobose, dehiscence longitudinal, borne on androgynophore; androgynophore fleshy, segmented, short, with 5 glands opposite to petals. Ovary 2-5-loculed; ovules 2 per locule; style simple; stigma 2-5-lobed. Fruit a capsule nearly globose, 3-6-valved, spiny or strigose, loculicidally dehiscent or indehiscent, spine tips pointed, straight or hooked. Seeds with endosperm; cotyledons fleshy, epigeous.
Ovary often tubercled or echinulate, each tubercle surmounted by one or more minute bristles, 2–5-locular with 2 pendulous collateral ovules in each loculus or falsely 10-locular by the intrusion of longitudinal false septa; style terete, about as long as the stamens; stigma entire or very shortly 2–5-lobed.
Stamens 4–40, raised on a short glabrous androgynophore or torus with a glandular patch just above each petal base; apex of the androgynophore produced into a ciliate or pubescent or villous disk or annulus between which and the ovary the stamens are inserted.
Fruit a capsule, dividing into 3–5-valves with 1–2 seeds per loculus or indehiscent, usually globose, sometimes ovoid, echinate or setose.
Seeds obovoid or subreniform; testa rather leathery and brown; embryo straight; cotyledons flat, suborbicular; endosperm fleshy, scanty.
Leaves alternate, petiolate, simple or digitate, often lobed, serrate or crenate, often several-nerved from the base; stipules lateral.
Petals 5, yellow or orange, linear to obovate, narrowed to the base, and often hairy at the base or just above it.
Sepals 5, usually linear and with a short horn just behind the apex, usually stellately hairy without.
Annual or perennial herbs, shrublets or shrubs, sometimes with annual stem from a woody rootstock.
Flowers actinomorphic, borne in cymes in terminal inflorescences or at the nodes.