Annual or perennial herbs or shrubs, glabrous, scabrous or pilose. Leaves alternate, opposite, rarely in whorls of 3, sessile or petiolate, entire or serrate. Inflorescence an indeterminate raceme or spike; flowers bisexual, 4-merous, single (rarely in fascicles) in axils of bracts with a pair of bracteoles (in G. nodulosus bracteoles replaced by 4 hairs). Sepals deltoid, lanceolate or cordate, often with prominent basal callus, persistent in fruit. Petals hooded, keeled, usually shortly clawed. Stamens usually twice number of sepals; anthers oblong, 4-locular; filaments short. Ovary ovoid to hemispherical, incompletely 4-locular, ribbed and ornamented, never winged; styles clavate; stigmas fimbriate, usually red. Seed 1, occupying entire fruit.