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Leaves digitately 3–9-foliolate or rarely simple and entire or lobed, very rarely pedate, margins variously toothed; stipules present.
Seed usually 1, usually with a distinct dorsal crest or crests and often ± rugose.– Corolla puberulous or pubescent and glandular21.
Flower-bud ± cylindric or lageniform, apex rounded and often ± inflated, ± constricted at or near the middle.
Disk of 4 rather fleshy truncate or conical glands, ± free from each other but ± adnate to the ovary.
Erect, prostrate or climbing perennial herbs or shrubs; tendrils opposite the leaves or absent.
Style simple, subulate; stigma minutely 2-fid, or subentire and minutely sub-capitate.
Anthers usually with ± straight filaments not bending noticeably over the gynoecium.
Petals cucullate at the apex, becoming deflexed after anthesis, very caducous.
Inflorescences of leaf-opposed or axillary corymbose cymes.
Calyx entire or ± 4-toothed.
Flowers 4-merous.
